1942. Evie Sinclair has enlisted in the ATS to serve her country. Her mother disapproves, wanting her to stay and help run the household, but her father supports her bid for independence. Though conditions are austere and the training is intense, Evie is proud to become a qualified gunner girl, part of the team operating the anti-aircraft batteries. When she meets RAF bomber pilot Matt Goodwin at a dance, it's not long before they fall in love. But then tragedy strikes. Matt is killed on a mission, leaving Evie bereft - and expecting a child.

Left homeless after the Exeter Blitz, May must find a job if she's to put a roof over her head and help support her two younger sisters. Taking a job as housekeeper to farmer George, May soon ends up getting stuck into more than just cleaning and cooking. The Ministry of Agriculture will close Fair Maids Farm if it doesn't meet their produce targets, but George refuses to heed their warnings. With only two reluctant Land Girls to help, May receives unexpected guidance from Dan, a neighbouring farmer, whose kindness gets tongues wagging in the village.

East London, 1940. At the outbreak of war, hopeless romantic Prue Carmichael and her sister must leave their rural parish behind when their father is posted to a church in Stepney. To Prue, the blitz-ravaged streets of London seem an unlikely place for love to bloom – until a chance encounter throws rakish engineer Jack Quinn into her path. But as their connection deepens, his troubled past begins to emerge, and Prue realises Jack has secrets to hide.

It’s 1918 in German-occupied Bruges, Belgium. With luck, Evelien will make it to the end of the war and be given what she was promised: a prized painting in exchange for safeguarding her employer’s possessions. Until then, Evelien knows to keep her head down and stay out of trouble. But life never goes to plan, especially in war. A member of the Resistance approaches Evelien: steal a list of names hidden in her employer’s home.

1939 - American heiress Vivi Miles falls for naval officer Nathan as soon as she arrives in England. And, under the threat of war, they marry in a whirlwind before he leaves to join his ship. When Nathan returns from Dunkirk injured, he is distant, aloof, and no longer the man Vivi fell in love with. But it’s not just because of his brutal experiences of war. Nathan has a secret and Vivi suspects it’s linked to the mysterious evacuee at the secluded house in the woods on his Kent estate.

June 1914. Edie Moore is a Governess for Lord and Lady Moreland, living comfortably at the grand Downland House in Sussex. But, wanting more from life, she flees to Littlehampton, where she spent many idyllic childhood holidays. Desperate for work, Edie begins working as a chambermaid at the prestigious Beach Hotel, despite the menial tasks being a far cry from her previous job. But as she navigates her new life, friendships, and rivalries, she also grows closer to charming, cheerful porter, Charlie Cobbett, and finally finds the happiness she has been searching for.

Staffordshire, 1911. Money is in short supply for Ginnie Jones’ family, and she is sent to Haddon Workhouse. When she meets Clara and Sam, she hopes that she has made friends for life...until tragedy strikes. Three years later, Ginnie finds work as a mouldrunner at the Potteries, but never stops thinking about her friends. When Sam and Ginnie are reunited, their bond is as strong as ever, until Sam is sent to fight in the war. Can Ginnie find the joy that she’s never had? 

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

When duty calls, East End girl Minnie Wolton isn’t afraid to do her bit. She’s excited to be joining the ATS girls and although she knows life at the barracks will be tough, she’s not one to quit! She just hopes the other girls are all good sorts. With her husband away fighting, Eileen Ruffel is glad of the break from her unhappy marriage. She just wants to make a success of the ATS girls and maybe find happiness too. Along with the other girls, Minnie and Eileen strike up a firm friendship. The training is hard, but they are determined to prove they are the best new recruits.

Lou Channer wants a life outside of North Devon, somewhere she's never left. She yearns to contribute to the war effort and takes a job as a clerk in the Royal Canadian Naval Yard in Plymouth, lodging with other girls from the depot who take her under their wing. When Lou catches the eye of local wheeler-dealer Harry, who dazzles her with nights about town, she finally feels like one of the girls. And when Lieutenant Douglas Ross asks her out, Lou can't decide who to give her heart to. But during war, tragedy is always just around the corner....

Pregnant, Mary Jane Starlings secret wedding to Paddy Redfern ends in disaster when her fiancé is murdered on the way to the church. Paddy’s wayward twin ‘Red’ intercepts Mary Jane and warns that they must flee Ireland for fear of reprisals from her family, never to return. Too young and terrified to question Red’s motives, Mary Jane, is hurriedly escorted to Liverpool where she quickly discovers Red’s good Samaritan act is a sham when he abandons her homeless and destitute.

1884 - When Betsan Morgan’s mother succumbs to a devastating illness, she doesn't think her life could get much worse. But then her father moves in his new beau, an unserious flitty woman and coincidentally also the bargirl in the pub he works in, and she soon learns her house is no longer a home. All she can do is sit in the attic and stare at the portrait of her mother. Her new stepmother, Elinor, couldn’t even let them have the grace of keeping it in the hallway.

Born to a French mother and an English father, Charlotte Bray has always felt like an outsider in her Cornish fishing village. She spends her days on her father's fishing boat, helping to feed a nation at war. But the war brings devastation, and her world is forever changed by tragedy. With nothing left to lose, she accepts a mysterious invitation to work for the War Office as a spy. She soon finds herself in Nazi-occupied Paris with a new identity. Charlotte begins working under the sombre but handsome Frenchman Pierre Lesieur. But do his allegiances lie elsewhere?

Cambridgeshire - March 1939. As the clouds of war begin to gather in Europe, the Talbot family of rural Blackberry Farm will be torn apart, just as so many families all over the world will be. Life will never be the same again. Whilst in London, the Salmons family will feel the pain of parting and loss. Brought together by war, the two families become intertwined and, as the outlook looks bleak, they must draw on each other’s strength to fight through the hard times.

Clara Newton is the new Housemother of Shilling Grange Orphanage. Many of the children have been bombed out of their homes and left without families, their lives torn apart by the war, just like Clara’s. Devastated by the loss of her fiancé, a brave American pilot, she is just looking for a place to start again. But the orphans are in desperate need of her help.