Lou Channer wants a life outside of North Devon, somewhere she's never left. She yearns to contribute to the war effort and takes a job as a clerk in the Royal Canadian Naval Yard in Plymouth, lodging with other girls from the depot who take her under their wing. When Lou catches the eye of local wheeler-dealer Harry, who dazzles her with nights about town, she finally feels like one of the girls. And when Lieutenant Douglas Ross asks her out, Lou can't decide who to give her heart to. But during war, tragedy is always just around the corner....

Left homeless after the Exeter Blitz, May must find a job if she's to put a roof over her head and help support her two younger sisters. Taking a job as housekeeper to farmer George, May soon ends up getting stuck into more than just cleaning and cooking. The Ministry of Agriculture will close Fair Maids Farm if it doesn't meet their produce targets, but George refuses to heed their warnings. With only two reluctant Land Girls to help, May receives unexpected guidance from Dan, a neighbouring farmer, whose kindness gets tongues wagging in the village.

Esme Colborne is about to marry Richard Trevannion. When she finds out she is adopted - from a working-class family - she cannot allow Richard to marry so far beneath his station. Fleeing the life she knew, she starts working as a ‘decoy woman’, testing British undercover operatives who may otherwise reveal secrets in a moment of weakness. However, she still has feelings for Richard. Will she be brave enough to risk having her heart broken all over again? 

October 1940. Twenty-three-year-old Bobby Bancroft is working as a typist for a city newspaper, but she longs to be breaking the news herself. When she successfully applies for a junior reporter role at The Tyke, a magazine serving the Yorkshire Dales, she's thrilled – even if she only gets the position because so many men are away fighting. Bobby moves to the sleepy village of Silverdale, but she quickly discovers life in the countryside is a different world.

It's 1943, and the Second World War is raging. Ruby Mottram works for her local newspaper, the Bartonford Herald, typing up adverts and obituaries, whilst dreaming of a more exciting life. Between her shifts as an ARP warden and caring for her ailing father, the chance for escape doesn't come often to Devon. Meanwhile, in America's Deep South, Sam Archer is hatching a plan to raise enough money to get his mother and sister away from his abusive stepfather. 

October 1940. When VAD nurse Evie narrowly escapes being killed in an air raid during the London Blitz, it propels her to make a life-changing decision to break free of her troubled and unhappy life. She escapes to the Norfolk countryside to start afresh, with a job at the newly opened Great Plumstead Hall hospital, and a wonderful new home at Rookery House. The community of Great Plumstead welcomes evacuees to the village—mothers and children bombed out of their London homes. Sisters Prue and Thea, along with members of The Mother's Day Club, help the new arrivals settle in.

East London, 1940. At the outbreak of war, hopeless romantic Prue Carmichael and her sister must leave their rural parish behind when their father is posted to a church in Stepney. To Prue, the blitz-ravaged streets of London seem an unlikely place for love to bloom – until a chance encounter throws rakish engineer Jack Quinn into her path. But as their connection deepens, his troubled past begins to emerge, and Prue realises Jack has secrets to hide.

As war rages, Barbara Sinclair is desperate to escape her unhappy home life. And with the threat of German bombs ever present, Barbara reluctantly agrees to marry John, her childhood friend, who is leaving to join the RAF. But an encounter with Alex Everton, a dashing Spitfire pilot, complicates matters for Barbara. With emotions running high, she begins to question whether she has made a terrible mistake. With the constant threat of death all around her, Barbara must try to find a way to deal with the complexities of her difficult home life and her emotional relationships, too.

1942. Evie Sinclair has enlisted in the ATS to serve her country. Her mother disapproves, wanting her to stay and help run the household, but her father supports her bid for independence. Though conditions are austere and the training is intense, Evie is proud to become a qualified gunner girl, part of the team operating the anti-aircraft batteries. When she meets RAF bomber pilot Matt Goodwin at a dance, it's not long before they fall in love. But then tragedy strikes. Matt is killed on a mission, leaving Evie bereft - and expecting a child.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

Jane’s parents think Eddie’s rebellious ways will lead their beloved daughter into trouble. They forbid her from seeing Eddie, telling Jane to get a proper job as a nurse. But Jane cannot stand the idea of life without the boy she loves and secretly continues to see Eddie. The day she finds out that Eddie has betrayed her, she is utterly distraught. Were her parents right all along?

1940, London. When Cordelia accepts the post of head librarian in Silver Town Library, her mother is more than a little disapproving. The East End has high levels of poverty and illiteracy, and her mother says it’s no place for a woman of her status. But Cordelia is determined to make a difference in these times of strife, and along with her colleagues, Jane and Mavis, she begins to help the local community, making sure everyone knows what the library can offer them. And maybe even a romance will blossom.

Against the chaotic backdrop of World War II, Grace Murphy is working as a midwife in Doncaster Royal Infirmary with her two friends, Clodagh and Patsy. In between delivering babies and tending wounded airmen, the girls endure bombing raids, rationing and the hardships and heartbreak that war brings. But in the faces of the new babies she welcomes to the world, Grace always looks for hope, as she dreams of a time when the war will end.

Essex 1941. Unhappy with life at home with her privileged family, Hannah Austen-Bagshaw runs away to join the Land Girls Army, carrying with her a dark and terrible secret. Life on the farm is tough, but when Hannah meets handsome working-class flight lieutenant Jack, they form an immediate friendship. But unbeknownst to Jack, Hannah is sheltering a young German pilot who has crash landed in a nearby field. Torn between her duty and her humanity, Hannah knows she risks being arrested as a traitor, but she cannot see the young pilot suffer.

Following a humiliating experience involving the man she thought she’d marry, Iris Tredwick signs up to the Wrens in order to escape and - hopefully - find ‘the right sort’ of man to please her mother. After a bumpy start, Iris manages to befriend outspoken Mary and dreamer Sally as they are sent to their first posting - in Orkney. There she meets engineer Rob, whose flirtatious nature both charms and confounds straight-laced Iris. Much more appropriate for her is local doctor Stewart, if only she felt the same spark for him as she does for Rob.