Cliffehaven, December 1946. From a hill above the town of Cliffehaven, a young woman makes a heart-breaking choice that will change the course of her life forever. Hours later, a baby is found in the Nativity crib of the local church. Who could have left him there, and why? The mystery preoccupies everyone in Cliffehaven - not least Peggy Reilly, who has enough to contend with at Beach View Boarding House without this extra secret to unravel.

When sixteen-year-old Sally is evacuated to the English south coast, she is terrified by what lies ahead of her. All she knows are the sights and sounds of London's East End – but Sally swallows her tears as they leave the familiar landmarks behind, knowing that she has to be a Grown-Up Girl and play mother to her six-year-old brother Ernie. Playing mother is nothing new for Sally – their real mother Florrie, a good-time girl, hasn't even come to the station to wave them off and Ernie, crippled at an early age by polio, is used to depending on his older sister.

Lancashire, 1895. When her controlling stepfather suddenly dies, it seems that Elinor Pendleton finally has a chance of freedom. But her hopes are soon dashed when she learns that the thuggish Jason Stafford has inherited every penny, and is determined to have Elinor too. Forced to flee with her maid, Maude, Elinor finds shelter with Maude's distant cousin in the remote village on the shore of Jubilee Lake. But Walter Crossley has troubles of his own. Having lost his closest family in a tragic accident, he needs one of his grandsons to return from America to inherit his farm.

In a village in Wiltshire, just off a busy road, Larch Tree Lane winds slowly up a low hill. Among the residents is Maggie Hatherall, full of life despite her advancing years, who worries what she is going to do with her 200-year-old home. Lucia Grey has escaped a bad marriage but her ex is reluctant to let her start a new life free of him. And for architect Corin Drayton, an unexpected inheritance is the start of a complicated family wrangle. From the cottages and farms to the local shop, pub and old manor house, the residents' lives are set to intertwine and relationships bloom.

As war looms in Europe, 18-year-old Jane Shaw runs the household as an unpaid servant on a meagre budget. When tragedy strikes and she loses her beloved younger brother in suspicious circumstances, Jane takes a position in service looking after Ned, a troubled young boy. Here she meets dashing David Heron, Ned's Uncle and they form an immediate attachment until she is dismissed after being accused of stealing a piece of jewellery. Returning home, Jane and her sister, Melia, are soon orphaned and left at the mercy of their Aunt Alice.

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.

Hampshire, 1913. Amidst the glitz and glamour of England’s High Society, Sarah Trenwith is more of a wallflower living in the shadow of her beautiful sister Marianne. But as the war approaches Sarah will have to step out of the place society and her family have assigned her if she’s to do her part and claim the love she always dreamed of from afar.

Wakefield 1870. On the banks of the River Calder, Meg Taylor struggles to care for her younger siblings and sick mother whilst her father is away working as a boatman on the canals. The slums where they live are rife with disease, and Meg longs for a different life, away from the grinding poverty, but she'll never leave her family. But with the canals slowly dying as trade moves to the railways, and with Meg’s father stubbornly holding onto the past, Meg fears her family’s future is going to get even bleaker. If only there was a way she could save them.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

When Emily Mattison falls victim to a near-fatal accident shortly after receiving an unexpected inheritance from a distant cousin, her ruthless nephew George seizes the opportunity to take control of his aunt's assets. It's only when Emily reaches The Drover's Hope, the former pub on the edge of the Lancashire moors bequeathed to her by her late cousin Penelope, that she begins to feel safe. She also discovers that love can be found in the most unexpected places. But it's not so easy to escape the clutches of someone as determined as George.

Bristol, 1939. School leaver Maisie Miles suspects her father, a small-time crook, has an ulterior motive for insisting she gets a job at the W. D. & H. O. Wills tobacco factory but keeps it to herself. She's befriended by effervescent Phyllis Mason and kind-hearted Bridget Milligan who take pity on her and take Maisie under their wing. But beneath their happy-go-lucky exteriors, they all harbour dreams and worries about what the future holds. 

At the heart of Dover's Market Square lies the Castle's Café. A bustling, homely place run by the formidable Nellie Castle and her six children. Since the scandalous birth of her beloved son 10 years before, Marianne, Nellie's eldest daughter, has preferred to stay in the kitchen, hidden away from the town gossips. Overcome with shame, Marianne has never revealed the name of Donny's father - not even to her own mother. But with World War II just around the corner, Marianne's past catches up with her. And suddenly the lives of the Castle family become a lot more complicated.

It's 1938, and Cadi is chosen to be Rose Queen in the annual Rose Fete. She is thrilled to be treated like royalty for the day. But deep down she is desperate to leave the Welsh mining community where she grew up. When war is declared, Cadi and her best friend, Poppy, see a chance to escape. Cadi leaves behind her parents and local boy Aled, whom she is meant to marry, for Liverpool.

When Mara Gregory receives a letter from the father whom she believed to have died when she was a child, her world is turned upside down. Aaron Buchanan only discovered that he had a daughter a couple of years ago and now he's desperate to play a part in her life. In the face of her mother's opposition, Mara arranges to meet her father and his family. In a breath-taking corner of the world, amid a waterfront community on Australia's west coast, will Mara find him the disappointment that her mother promises?

October 1940. When VAD nurse Evie narrowly escapes being killed in an air raid during the London Blitz, it propels her to make a life-changing decision to break free of her troubled and unhappy life. She escapes to the Norfolk countryside to start afresh, with a job at the newly opened Great Plumstead Hall hospital, and a wonderful new home at Rookery House. The community of Great Plumstead welcomes evacuees to the village—mothers and children bombed out of their London homes. Sisters Prue and Thea, along with members of The Mother's Day Club, help the new arrivals settle in.