October 1940. When VAD nurse Evie narrowly escapes being killed in an air raid during the London Blitz, it propels her to make a life-changing decision to break free of her troubled and unhappy life. She escapes to the Norfolk countryside to start afresh, with a job at the newly opened Great Plumstead Hall hospital, and a wonderful new home at Rookery House. The community of Great Plumstead welcomes evacuees to the village—mothers and children bombed out of their London homes. Sisters Prue and Thea, along with members of The Mother's Day Club, help the new arrivals settle in.

As war rages, Barbara Sinclair is desperate to escape her unhappy home life. And with the threat of German bombs ever present, Barbara reluctantly agrees to marry John, her childhood friend, who is leaving to join the RAF. But an encounter with Alex Everton, a dashing Spitfire pilot, complicates matters for Barbara. With emotions running high, she begins to question whether she has made a terrible mistake. With the constant threat of death all around her, Barbara must try to find a way to deal with the complexities of her difficult home life and her emotional relationships, too.

August 1940: As the Luftwaffe swarm over Liverpool, Shane Quinn decides to move his family back to the safety of Ireland. But his only child, the beautiful Dana, would rather stay and serve her country than flee to a foreign land. Determined to make it on her own, she joins the WAAF with newfound pals Patty and Lucy. There’s plenty of excitement to be had on a RAF station, even a chance or two at love.

In February, 1922, at the western-most entrance to Victoria Station in Manchester, a massive plaque was unveiled. Beneath a vast tiled map showing the lines of the Lancashire and Yorkshire Railway network, a series of seven bronze panels recorded the names of the men of the Lancashire and Yorkshire Railway who gave their lives for King and Country in the Great War - a total of 1,460 names. In March, 1940, a group of women of varying ages and backgrounds stands in front of the memorial, ready to do its bit in this new World War.... 

It's 1939, and the Brogan family of London's East End are ready to show Hitler what for. But things don't seem so rosy when rationing, evacuation and air raids start to put this larger-than-life family to the test. When a mysterious young man arrives at the Brogans' local parish church, he provides just the dazzling distraction they need - and, for eldest daughter Mattie, the promise of more than she'd ever wished for. But as the pair fall deeper in love, they are drawn into secret dangers, rife on the very London streets they call home.

Wakefield 1870. On the banks of the River Calder, Meg Taylor struggles to care for her younger siblings and sick mother whilst her father is away working as a boatman on the canals. The slums where they live are rife with disease, and Meg longs for a different life, away from the grinding poverty, but she'll never leave her family. But with the canals slowly dying as trade moves to the railways, and with Meg’s father stubbornly holding onto the past, Meg fears her family’s future is going to get even bleaker. If only there was a way she could save them.

It is 1937, and Alice, Edith and Marg continue to face hardships every day, growing up on one of the poorest streets in Blackpool. Penniless, their friendship has helped them survive this far, but it'll take more than that to see them through the dark days ahead. Alice is coping with a violent father. Marg is left reeling after a dark secret about her birth comes to light and threatens to destroy the life she knows, and Edith is fighting to protect her alcoholic mother from the shame of their neighbours and keep her brother on the straight and narrow. 

1944. With deadly doodlebugs terrorising London, Bessie Rushbrook agrees to give shelter to eight-year-old evacuee Marigold in her home. But, the little girl is no stranger, and Bessie must honour the promise she gave to Marigold's mother and not reveal her connection to the child. Marigold's arrival stirs up Bessie's memories from the past when the world was at war the first time. She is forced to face her actions from those days and question the secret that she's long kept hidden. When Grace is injured in London, Bessie makes the heart-wrenching decision to confess her secret.

Blackpool, 1893. Tilly has come a long way from the run-down tenements in which she grew up. She has a small but comfortable home, a loving, handsome husband, two beautiful little'uns - Babs and Beth - and she earns herself a little money weaving wicker baskets. Life is good. Until the day Tilly returns home to find a policeman standing on her doorstep. Her Arthur won't be coming home tonight - nor any night - having fallen to his death whilst working on Blackpool tower. Suddenly Tilly is her daughters' sole protector, and she's never felt more alone.

London, 1939. When tragedy strikes, Libby Andrews is left homeless and destitute, fending for herself on the capital's most dangerous streets. Desperate times call for desperate measures, and Libby is on the cusp of making a decision that could jeopardise her future when a local boy saves her from ruin. The kindness of this stranger sets her on a different path, and Libby heads to Liverpool with a determined mind and hope in her heart. There, she reconnects with long lost family - but will she be able to uncover the truth that tore them apart all those years ago?

Sunderland, 1940, and the women go to the shipyards to do their bit for the war effort. Polly never dreamed she would be able to work in the shipyards like the men in her family but times are tough and her new job ends up giving her more than she ever expected when she meets enigmatic dock diver Tommy Watts. During the day, head welder Rosie teaches her fledgling flock of trainees their new trade, but at night she hides a secret life. And mother hen Gloria signs up to escape her brutal husband, but finds she cannot run....

1940, Liverpool: Alice Turner and her best friend, Millie Markham, work for the war effort at Rootes munitions factory, making shell caps and Halifax bombers. Alice’s sweetheart, Terry, is home from the front for a brief period of leave. But the honeymoon is soon over. Terry must go back to the dangers of Nazi-occupied France, and the ever-present air raid sirens quickly bring Alice back down to earth. 

Bristol 1939. At forty years of age, Mary Anne Randall still turns heads. With an abusive husband spending most of his wages on beer, she has resigned herself to be the sole breadwinner to protect her family and keep the wolves from the door. In order to make ends meet, Mary Anne runs a pawnbroking business from the wash house at the end of the garden. Following the outbreak of war an opportunity presents itself to escape her loveless marriage and find true love. Will she take that chance, or carry on living just for her children and not for herself?

Laura Collins is struggling to make a good life for her daughters, Tess and Tina. So when she is offered a good job, with accommodation, she is delighted. Life is very different for the Brewsters who live on Manor Farm in Herefordshire. Danny plans to marry one day so that he can take over the farm, whilst his cousin, Phil, longs to escape the world of farming as soon as possible. War changes all their plans.

Hull, 1940. Widowed Edie Copeland has just arrived on Exchange Street, heavily pregnant and with heavy suitcase in tow. She left York after tragedy struck to make a fresh start, but she's a stranger to this street, and her new landlord doesn't hesitate to remind her of this, widow or not. Luckily, the neighbours are a little more welcoming and Edie is soon made to feel at home by the Maguires and the Scuttles. As air raids sound and the war feels closer than ever, the community has to stick together. But Edie is hiding something, and she doesn't know how much longer her secret is safe.