The three sisters of Victory Walk in East London will face heartache and tears on the home front. As rationing, blackouts and bombs start to bite, life will change forever for the three sisters.

When duty calls, East End girl Minnie Wolton isn’t afraid to do her bit. She’s excited to be joining the ATS girls and although she knows life at the barracks will be tough, she’s not one to quit! She just hopes the other girls are all good sorts. With her husband away fighting, Eileen Ruffel is glad of the break from her unhappy marriage. She just wants to make a success of the ATS girls and maybe find happiness too. Along with the other girls, Minnie and Eileen strike up a firm friendship. The training is hard, but they are determined to prove they are the best new recruits.

In a village in Wiltshire, just off a busy road, Larch Tree Lane winds slowly up a low hill. Among the residents is Maggie Hatherall, full of life despite her advancing years, who worries what she is going to do with her 200-year-old home. Lucia Grey has escaped a bad marriage but her ex is reluctant to let her start a new life free of him. And for architect Corin Drayton, an unexpected inheritance is the start of a complicated family wrangle. From the cottages and farms to the local shop, pub and old manor house, the residents' lives are set to intertwine and relationships bloom.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

Autumn 1941, and as female conscription looms, three women arrive at the Bell Works Factory in Hucknall to work on the Spitfire production line. Lily soon realises her real dream is to pilot the planes; Edna longs to be reunited with her evacuee daughter, but soon discovers she has a much bigger problem; and Jeannie fears the worst after weeks of no news from her fiancé. Despite all the heartache, with a bit of festive cheer and 'can do' attitude, The Engine Girls will pull together and do all they can to ensure Britain's safe for the New Year.

Cambridgeshire - March 1939. As the clouds of war begin to gather in Europe, the Talbot family of rural Blackberry Farm will be torn apart, just as so many families all over the world will be. Life will never be the same again. Whilst in London, the Salmons family will feel the pain of parting and loss. Brought together by war, the two families become intertwined and, as the outlook looks bleak, they must draw on each other’s strength to fight through the hard times.

Hampshire, 1913. Amidst the glitz and glamour of England’s High Society, Sarah Trenwith is more of a wallflower living in the shadow of her beautiful sister Marianne. But as the war approaches Sarah will have to step out of the place society and her family have assigned her if she’s to do her part and claim the love she always dreamed of from afar.

September 1939. London is in blackout; war has been declared, but Dulcie Treadwell can think only of her heartbreak over American broadcaster, Glenn Reeves, who didn’t say goodbye before leaving for Berlin. Posted to Bletchley Park, Dulcie must concentrate on helping the war effort by cracking the German Enigma codes. The hours are long and the conditions tough, with little recognition from above. Until she breaks her first code. Dulcie finally settles into something she was born to do. But just as she’s proving her worth, a shocking betrayal leads to Dulcie’s brutal dismissal.

As war rages, Barbara Sinclair is desperate to escape her unhappy home life. And with the threat of German bombs ever present, Barbara reluctantly agrees to marry John, her childhood friend, who is leaving to join the RAF. But an encounter with Alex Everton, a dashing Spitfire pilot, complicates matters for Barbara. With emotions running high, she begins to question whether she has made a terrible mistake. With the constant threat of death all around her, Barbara must try to find a way to deal with the complexities of her difficult home life and her emotional relationships, too.

It's 1944, and Rosie Sinclair is full of pride to be doing her bit for the war effort as a driver at Cottisbourne Park - the secret heart of Britain's fight against Germany, where a team of brilliant and eccentric codebreakers are battling to save the country. But when she's given a new mission to drive Major-General 'Bluff' Kingsley-Flynn down to Cottisbourne, Rosie finds herself on the frontline of a new battle - to uncover a possible spy at the Park who is jeopardising their vital work, and to resist her own growing attraction to the dashing Bluff himself.... 

1941. The Blitz rages over London. And even in Cornwall, the war is being fought. When Violet loses her sister in the Blitz, she must take her nieces to safety in Cornwall. On the coast, she meets carefree chorus girl Eva, who is also running from the dangers of London. But Porthcurno hides a secret military base, and soon Violet and Eva realise there’s a battle to fight in Cornwall, too. Together with local Hazel, who works on the base, they must come together to help the war effort. But will their friendship be enough to keep them safe? 

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.

Bristol, 1939. School leaver Maisie Miles suspects her father, a small-time crook, has an ulterior motive for insisting she gets a job at the W. D. & H. O. Wills tobacco factory but keeps it to herself. She's befriended by effervescent Phyllis Mason and kind-hearted Bridget Milligan who take pity on her and take Maisie under their wing. But beneath their happy-go-lucky exteriors, they all harbour dreams and worries about what the future holds. 

In the heart of Dressmakers' Alley, Madame Pauline’s sweatshop is rumoured to be a particularly unpleasant place to work. Filled with seamstresses paid a pittance who turn a blind eye to what darker activities are hidden behind locked doors. Young Winnie Brown is keen to prove her value to the Women Movement and secures a job as a seamstress to investigate the whispers of some unsavoury goings on. Her concerns are soon justified when she hears a desperate cry for help.

In February, 1922, at the western-most entrance to Victoria Station in Manchester, a massive plaque was unveiled. Beneath a vast tiled map showing the lines of the Lancashire and Yorkshire Railway network, a series of seven bronze panels recorded the names of the men of the Lancashire and Yorkshire Railway who gave their lives for King and Country in the Great War - a total of 1,460 names. In March, 1940, a group of women of varying ages and backgrounds stands in front of the memorial, ready to do its bit in this new World War....