It is 1937, and Alice, Edith and Marg continue to face hardships every day, growing up on one of the poorest streets in Blackpool. Penniless, their friendship has helped them survive this far, but it'll take more than that to see them through the dark days ahead. Alice is coping with a violent father. Marg is left reeling after a dark secret about her birth comes to light and threatens to destroy the life she knows, and Edith is fighting to protect her alcoholic mother from the shame of their neighbours and keep her brother on the straight and narrow. 

As war looms in Europe, 18-year-old Jane Shaw runs the household as an unpaid servant on a meagre budget. When tragedy strikes and she loses her beloved younger brother in suspicious circumstances, Jane takes a position in service looking after Ned, a troubled young boy. Here she meets dashing David Heron, Ned's Uncle and they form an immediate attachment until she is dismissed after being accused of stealing a piece of jewellery. Returning home, Jane and her sister, Melia, are soon orphaned and left at the mercy of their Aunt Alice.

Bristol, 1939. School leaver Maisie Miles suspects her father, a small-time crook, has an ulterior motive for insisting she gets a job at the W. D. & H. O. Wills tobacco factory but keeps it to herself. She's befriended by effervescent Phyllis Mason and kind-hearted Bridget Milligan who take pity on her and take Maisie under their wing. But beneath their happy-go-lucky exteriors, they all harbour dreams and worries about what the future holds. 

Lady Joanna Harcourt and her daughter, Sarah, discover their life of luxury at Goodwill House is over. Because with Lord Harcourt away fighting, the Harcourt girls are facing financial ruin. Lady Joanna is terrified of losing their home, but for Sarah, this means her dreams of becoming a doctor are over. Headstrong Sarah isn’t one to quit! War or not, she’s determined to find a way to save her home and follow her dreams - whatever it takes.

When duty calls, East End girl Minnie Wolton isn’t afraid to do her bit. She’s excited to be joining the ATS girls and although she knows life at the barracks will be tough, she’s not one to quit! She just hopes the other girls are all good sorts. With her husband away fighting, Eileen Ruffel is glad of the break from her unhappy marriage. She just wants to make a success of the ATS girls and maybe find happiness too. Along with the other girls, Minnie and Eileen strike up a firm friendship. The training is hard, but they are determined to prove they are the best new recruits.

1940, London. When Cordelia accepts the post of head librarian in Silver Town Library, her mother is more than a little disapproving. The East End has high levels of poverty and illiteracy, and her mother says it’s no place for a woman of her status. But Cordelia is determined to make a difference in these times of strife, and along with her colleagues, Jane and Mavis, she begins to help the local community, making sure everyone knows what the library can offer them. And maybe even a romance will blossom.

Following a humiliating experience involving the man she thought she’d marry, Iris Tredwick signs up to the Wrens in order to escape and - hopefully - find ‘the right sort’ of man to please her mother. After a bumpy start, Iris manages to befriend outspoken Mary and dreamer Sally as they are sent to their first posting - in Orkney. There she meets engineer Rob, whose flirtatious nature both charms and confounds straight-laced Iris. Much more appropriate for her is local doctor Stewart, if only she felt the same spark for him as she does for Rob.

Flying instructor Ellie Simpson has grown up a tomboy. She's never had an interest in the latest fashions or finding a husband; her only passion is flying her beloved Tiger Moth. But when war is declared, she is no longer be able to do what she loves most - fly.

Cambridgeshire - March 1939. As the clouds of war begin to gather in Europe, the Talbot family of rural Blackberry Farm will be torn apart, just as so many families all over the world will be. Life will never be the same again. Whilst in London, the Salmons family will feel the pain of parting and loss. Brought together by war, the two families become intertwined and, as the outlook looks bleak, they must draw on each other’s strength to fight through the hard times.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

Kitty Wilson, who, after losing her father, rejects the unsuitable suitor her mother has chosen. Abandoned by her mother, Kitty faces the challenges of navigating life alone. Amidst the turmoil, Kitty finds solace and support from friends, securing a promotion at Harpers Emporium. However, tragedy looms, threatening to shatter her newfound stability. Sally and Ben Harper face their own challenges. A decision by Ben regarding the restaurant, made without consulting his partner Jack Burrows, jeopardizes not only their friendship but also the stability of Harpers itself.

East London, 1940. At the outbreak of war, hopeless romantic Prue Carmichael and her sister must leave their rural parish behind when their father is posted to a church in Stepney. To Prue, the blitz-ravaged streets of London seem an unlikely place for love to bloom – until a chance encounter throws rakish engineer Jack Quinn into her path. But as their connection deepens, his troubled past begins to emerge, and Prue realises Jack has secrets to hide.

October 1940. When VAD nurse Evie narrowly escapes being killed in an air raid during the London Blitz, it propels her to make a life-changing decision to break free of her troubled and unhappy life. She escapes to the Norfolk countryside to start afresh, with a job at the newly opened Great Plumstead Hall hospital, and a wonderful new home at Rookery House. The community of Great Plumstead welcomes evacuees to the village—mothers and children bombed out of their London homes. Sisters Prue and Thea, along with members of The Mother's Day Club, help the new arrivals settle in.

At the heart of Dover's Market Square lies the Castle's Café. A bustling, homely place run by the formidable Nellie Castle and her six children. Since the scandalous birth of her beloved son 10 years before, Marianne, Nellie's eldest daughter, has preferred to stay in the kitchen, hidden away from the town gossips. Overcome with shame, Marianne has never revealed the name of Donny's father - not even to her own mother. But with World War II just around the corner, Marianne's past catches up with her. And suddenly the lives of the Castle family become a lot more complicated.

As war rages, Barbara Sinclair is desperate to escape her unhappy home life. And with the threat of German bombs ever present, Barbara reluctantly agrees to marry John, her childhood friend, who is leaving to join the RAF. But an encounter with Alex Everton, a dashing Spitfire pilot, complicates matters for Barbara. With emotions running high, she begins to question whether she has made a terrible mistake. With the constant threat of death all around her, Barbara must try to find a way to deal with the complexities of her difficult home life and her emotional relationships, too.