Dilly is devastated: with her husband unable to work and four children already at home, they cannot afford to feed their newborn baby. Heartbroken, she delivers her baby girl to the Farthing family, who have just lost their own daughter to measles. This act of desperation will change the lives of both families irrevocably. But when the chance for true love finally comes, will Dilly choose family over her own happiness?

1850. When Opal Sharp finds herself and her younger siblings suddenly orphaned and destitute, she thinks things can get no worse. But soon three of them - including Opal - are struck down with the illness that took their father from them, and her brother Charlie is forced to make an impossible decision. Unable to afford a doctor, he knows the younger children will not survive. So, unbeknownst to Opal, Charlie takes their younger siblings to the workhouse, where he knows they will at least be fed and have a roof over their heads. Opal is heartbroken and struggles to forgive him. 

Dotty, Lucy and Annabelle all turn up for work at Coventry's department store Owen Owen at the same time as war is declared. Dotty is an orphan, Lucy is the sole carer for her little sister, and although Annabelle has led a life of privilege until now, everyone's having to pinch the pennies at the moment. The three form an unlikely bond, and with disaster at every turn, they're going to need each other...

On the outskirts of Nuneaton stands Hatters Hall, home to the mentally insane. One rainy, dismal day in 1857, Maria Mundy arrives there to work. She is ordered to care for Isabelle Montgomery, daughter of an influential land-owner, but it becomes apparent that she has been falsely imprisoned. It’s going to take courage, determination, and resolve - but freedom is always worth fighting for.

Warwickshire, 1840s: boys are destined for the pit and girls for the mill. Despite this, Maryann is happy there - until her mother dies. Her family collapses, leaving Maryann exhausted, and Toby, the boy she is set on marrying, insists they wait. Then Maryann is offered a position as nanny to the daughter of the mill owner, Wesley Marshall, who Maryann grows unexpectedly close to. But their relationship doesn’t go unnoticed, threatening to unleash a world of problems on them all....

Abandoned by their parents and left to fight for themselves, headstrong Rosalind Carey has no choice but to take charge of her younger brothers and sister in Devonshire’s grand Rockwood Castle. But their once much-loved home is crumbling like the family that lives within it. Living hand to mouth and desperate to provide for the estate that depends on them, the Carey family are one debt away from ruin. Until the day comes when the dashing Piers Blanchard appears on their doorstep from Cornwall, claiming he is Rosalind’s distant cousin and that Rockwood Castle is his.

London, 1854: 20-year-old Essie Chapman lives with her father in poverty-stricken Limehouse, working on the river as a boatman. Her life seems set before her, never to leave this part of London and forever at her father's beck and call. Then, one night, she must transport a mysterious man from a foreign ship to the banks of the Thames, a man who ends up renting a room in Essie's house, identifying himself only as 'Raven'.

In a village in Wiltshire, just off a busy road, Larch Tree Lane winds slowly up a low hill. Among the residents is Maggie Hatherall, full of life despite her advancing years, who worries what she is going to do with her 200-year-old home. Lucia Grey has escaped a bad marriage but her ex is reluctant to let her start a new life free of him. And for architect Corin Drayton, an unexpected inheritance is the start of a complicated family wrangle. From the cottages and farms to the local shop, pub and old manor house, the residents' lives are set to intertwine and relationships bloom.

When Emily Mattison falls victim to a near-fatal accident shortly after receiving an unexpected inheritance from a distant cousin, her ruthless nephew George seizes the opportunity to take control of his aunt's assets. It's only when Emily reaches The Drover's Hope, the former pub on the edge of the Lancashire moors bequeathed to her by her late cousin Penelope, that she begins to feel safe. She also discovers that love can be found in the most unexpected places. But it's not so easy to escape the clutches of someone as determined as George.

Lancashire, 1895. When her controlling stepfather suddenly dies, it seems that Elinor Pendleton finally has a chance of freedom. But her hopes are soon dashed when she learns that the thuggish Jason Stafford has inherited every penny, and is determined to have Elinor too. Forced to flee with her maid, Maude, Elinor finds shelter with Maude's distant cousin in the remote village on the shore of Jubilee Lake. But Walter Crossley has troubles of his own. Having lost his closest family in a tragic accident, he needs one of his grandsons to return from America to inherit his farm.

In February, 1922, at the western-most entrance to Victoria Station in Manchester, a massive plaque was unveiled. Beneath a vast tiled map showing the lines of the Lancashire and Yorkshire Railway network, a series of seven bronze panels recorded the names of the men of the Lancashire and Yorkshire Railway who gave their lives for King and Country in the Great War - a total of 1,460 names. In March, 1940, a group of women of varying ages and backgrounds stands in front of the memorial, ready to do its bit in this new World War.... 

Bristol, 1939. School leaver Maisie Miles suspects her father, a small-time crook, has an ulterior motive for insisting she gets a job at the W. D. & H. O. Wills tobacco factory but keeps it to herself. She's befriended by effervescent Phyllis Mason and kind-hearted Bridget Milligan who take pity on her and take Maisie under their wing. But beneath their happy-go-lucky exteriors, they all harbour dreams and worries about what the future holds. 

As war looms in Europe, 18-year-old Jane Shaw runs the household as an unpaid servant on a meagre budget. When tragedy strikes and she loses her beloved younger brother in suspicious circumstances, Jane takes a position in service looking after Ned, a troubled young boy. Here she meets dashing David Heron, Ned's Uncle and they form an immediate attachment until she is dismissed after being accused of stealing a piece of jewellery. Returning home, Jane and her sister, Melia, are soon orphaned and left at the mercy of their Aunt Alice.

It's 1939, and the Brogan family of London's East End are ready to show Hitler what for. But things don't seem so rosy when rationing, evacuation and air raids start to put this larger-than-life family to the test. When a mysterious young man arrives at the Brogans' local parish church, he provides just the dazzling distraction they need - and, for eldest daughter Mattie, the promise of more than she'd ever wished for. But as the pair fall deeper in love, they are drawn into secret dangers, rife on the very London streets they call home.

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.