London, September 1666. The Great Fire rages through the city, consuming everything in its path. Even the impregnable cathedral of St. Paul's is engulfed in flames and reduced to ruins. Among the crowds watching its destruction is James Marwood, son of a disgraced printer and reluctant government informer. In the aftermath of the fire, a semi-mummified body is discovered in the ashes of St. Paul's, in a tomb that should have been empty. The man's body has been mutilated, and his thumbs have been tied behind his back.

London, 1654. Oliver Cromwell is at the height of his power and has declared himself Lord Protector. Yet he has many enemies at home and abroad. London is a complex web of spies and merchants, priests and soldiers, exiles and assassins. One of the web's most fearsome spiders is Damian Seeker, agent of the Lord Protector. No one knows where Seeker comes from, who his family is, or even his real name. All that is known of him for certain is that he is utterly loyal to Cromwell and that nothing can be long hidden from him.

England, 1176. Aelred Barling, esteemed clerk to the justices of King Henry II, is dispatched from the royal court with his young assistant, Hugo Stanton, to investigate a brutal murder in a village outside York. The case appears straightforward. A suspect is under lock and key in the local prison, and the angry villagers are demanding swift justice. But when more bodies are discovered, certainty turns to doubt - and amid the chaos it becomes clear that nobody is above suspicion.

After Culloden, Iain MacGillivray was left for dead on Drumossie Moor. Wounded, his face brutally slashed, he survived only by pretending to be dead as the Redcoats patrolled the corpses of his Jacobite comrades. Six years later, with the clan chiefs routed and the Highlands subsumed into the British state, Iain lives a quiet life, working as a bookseller in Inverness. One day, after helping several of his regular customers, he notices a stranger lurking in the upper gallery of his shop, poring over his collection. But the man refuses to say what he's searching for.

1715. Jonas Flynt, ex-soldier and reluctant member of the Company of Rogues, a shady intelligence group run by ruthless spymaster Nathaniel Charters, is ordered to recover a missing document. Its contents could prove devastating in the wrong hands. On her deathbed, the late Queen Anne may have promised the nation to her half-brother James, the Old Pretender, rather than the new king, George I. But the will has been lost. It may decide the fate of the nation.