Blackpool, 1893. Tilly has come a long way from the run-down tenements in which she grew up. She has a small but comfortable home, a loving, handsome husband, two beautiful little'uns - Babs and Beth - and she earns herself a little money weaving wicker baskets. Life is good. Until the day Tilly returns home to find a policeman standing on her doorstep. Her Arthur won't be coming home tonight - nor any night - having fallen to his death whilst working on Blackpool tower. Suddenly Tilly is her daughters' sole protector, and she's never felt more alone.

It is 1937, and Alice, Edith and Marg continue to face hardships every day, growing up on one of the poorest streets in Blackpool. Penniless, their friendship has helped them survive this far, but it'll take more than that to see them through the dark days ahead. Alice is coping with a violent father. Marg is left reeling after a dark secret about her birth comes to light and threatens to destroy the life she knows, and Edith is fighting to protect her alcoholic mother from the shame of their neighbours and keep her brother on the straight and narrow. 

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.

1940, Liverpool: Alice Turner and her best friend, Millie Markham, work for the war effort at Rootes munitions factory, making shell caps and Halifax bombers. Alice’s sweetheart, Terry, is home from the front for a brief period of leave. But the honeymoon is soon over. Terry must go back to the dangers of Nazi-occupied France, and the ever-present air raid sirens quickly bring Alice back down to earth. 

August 1940: As the Luftwaffe swarm over Liverpool, Shane Quinn decides to move his family back to the safety of Ireland. But his only child, the beautiful Dana, would rather stay and serve her country than flee to a foreign land. Determined to make it on her own, she joins the WAAF with newfound pals Patty and Lucy. There’s plenty of excitement to be had on a RAF station, even a chance or two at love.

Life for Elsie is difficult as she struggles to cope with her alcoholic mother. Caring for her siblings and working long hours at Swift's jam factory in London’s Bermondsey is exhausting. Thankfully her lifelong friendship with Dot helps to smooth over life’s rough edges. When Elsie and Dot meet Millie Swift, they are nervous to be in the presence of the bosses’ daughter. Over time, they are surprised to feel so drawn to her, but should two East End girls be socialising in such circles?

After the hard work of being a maid at Wallington Manor in the lead-up to Christmas, Annie is very excited at the prospect of going home to Bethnal Green. She has missed her family, but the money she earns keeps them all afloat. Harriet is from the island of Guernsey and is visiting her aunt at Wallington Manor. When she has to leave for London Annie is asked to look after her. On the train journey a friendship blossoms. And when an accident happens their friendship is made even stronger. A friendship that would see both girls through pain, happiness, marriage and death.

Hull, 1940. Widowed Edie Copeland has just arrived on Exchange Street, heavily pregnant and with heavy suitcase in tow. She left York after tragedy struck to make a fresh start, but she's a stranger to this street, and her new landlord doesn't hesitate to remind her of this, widow or not. Luckily, the neighbours are a little more welcoming and Edie is soon made to feel at home by the Maguires and the Scuttles. As air raids sound and the war feels closer than ever, the community has to stick together. But Edie is hiding something, and she doesn't know how much longer her secret is safe.

In February, 1922, at the western-most entrance to Victoria Station in Manchester, a massive plaque was unveiled. Beneath a vast tiled map showing the lines of the Lancashire and Yorkshire Railway network, a series of seven bronze panels recorded the names of the men of the Lancashire and Yorkshire Railway who gave their lives for King and Country in the Great War - a total of 1,460 names. In March, 1940, a group of women of varying ages and backgrounds stands in front of the memorial, ready to do its bit in this new World War.... 

It wasn't a privileged childhood, but it was a happy one. Sybil and Lizzie Cream, brought up in a fisherman's cottage on the edge of the cold North Sea, were content to leave privilege where it belonged: with their friends the Wintertons. Christina Winterton was the same age as Sybil, and the two girls were inseparable, but it was Lizzie whom Ralph Winterton, three years older, found irresistible. Then war came to East Anglia, and so did Manchester-born Fenn Kitzmann, now of the American Army Air Force.

October 1940. When VAD nurse Evie narrowly escapes being killed in an air raid during the London Blitz, it propels her to make a life-changing decision to break free of her troubled and unhappy life. She escapes to the Norfolk countryside to start afresh, with a job at the newly opened Great Plumstead Hall hospital, and a wonderful new home at Rookery House. The community of Great Plumstead welcomes evacuees to the village—mothers and children bombed out of their London homes. Sisters Prue and Thea, along with members of The Mother's Day Club, help the new arrivals settle in.

Lancashire, 1895. When her controlling stepfather suddenly dies, it seems that Elinor Pendleton finally has a chance of freedom. But her hopes are soon dashed when she learns that the thuggish Jason Stafford has inherited every penny, and is determined to have Elinor too. Forced to flee with her maid, Maude, Elinor finds shelter with Maude's distant cousin in the remote village on the shore of Jubilee Lake. But Walter Crossley has troubles of his own. Having lost his closest family in a tragic accident, he needs one of his grandsons to return from America to inherit his farm.

As war rages, Barbara Sinclair is desperate to escape her unhappy home life. And with the threat of German bombs ever present, Barbara reluctantly agrees to marry John, her childhood friend, who is leaving to join the RAF. But an encounter with Alex Everton, a dashing Spitfire pilot, complicates matters for Barbara. With emotions running high, she begins to question whether she has made a terrible mistake. With the constant threat of death all around her, Barbara must try to find a way to deal with the complexities of her difficult home life and her emotional relationships, too.

Dympna Byrne lives a simple life with her family on the Connemara coast. She adores her father Micheál, and does her best to help her family by working hard and expecting little. But beneath the surface there are hidden secrets. Dympna’s mother idolises her clever eldest son, yet her attitude to Dympna is puzzling. So when her family find themselves in desperate need of money, it is Dympna who crosses the water to Liverpool to seek a better life. 

Living on the edge of the beautiful Norfolk Broads with her father, the only ripple in Tess Delamere's calm life is the disturbing dream about her dead mother which haunts her. She yearns to know more, but the arrival of a new stepmother heralds the end of Tess' hopes that her father might divulge the past. As she grows up, Tess slowly starts to put together the pieces herself. But the outbreak of war brings tragedy and upheaval, changing Tess' priorities....