When duty calls, East End girl Minnie Wolton isn’t afraid to do her bit. She’s excited to be joining the ATS girls and although she knows life at the barracks will be tough, she’s not one to quit! She just hopes the other girls are all good sorts. With her husband away fighting, Eileen Ruffel is glad of the break from her unhappy marriage. She just wants to make a success of the ATS girls and maybe find happiness too. Along with the other girls, Minnie and Eileen strike up a firm friendship. The training is hard, but they are determined to prove they are the best new recruits.

As war rages, Barbara Sinclair is desperate to escape her unhappy home life. And with the threat of German bombs ever present, Barbara reluctantly agrees to marry John, her childhood friend, who is leaving to join the RAF. But an encounter with Alex Everton, a dashing Spitfire pilot, complicates matters for Barbara. With emotions running high, she begins to question whether she has made a terrible mistake. With the constant threat of death all around her, Barbara must try to find a way to deal with the complexities of her difficult home life and her emotional relationships, too.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

Born in the workhouse and adopted by a former suffragette, Frances Brakespeare was encouraged from an early age to be strong, independent and to pursue a career as a doctor. The tragic loss of the love of her life in the Great War propels Frances to commit to her one true vocation. Rebelling against the unfair treatment of female doctors Frances is dismissed from her London post and things continue to take a turn for the worse when Izzy, her benefactress dies and Frances finds herself homeless.

1940, London. When Cordelia accepts the post of head librarian in Silver Town Library, her mother is more than a little disapproving. The East End has high levels of poverty and illiteracy, and her mother says it’s no place for a woman of her status. But Cordelia is determined to make a difference in these times of strife, and along with her colleagues, Jane and Mavis, she begins to help the local community, making sure everyone knows what the library can offer them. And maybe even a romance will blossom.

In February, 1922, at the western-most entrance to Victoria Station in Manchester, a massive plaque was unveiled. Beneath a vast tiled map showing the lines of the Lancashire and Yorkshire Railway network, a series of seven bronze panels recorded the names of the men of the Lancashire and Yorkshire Railway who gave their lives for King and Country in the Great War - a total of 1,460 names. In March, 1940, a group of women of varying ages and backgrounds stands in front of the memorial, ready to do its bit in this new World War.... 

1942. Evie Sinclair has enlisted in the ATS to serve her country. Her mother disapproves, wanting her to stay and help run the household, but her father supports her bid for independence. Though conditions are austere and the training is intense, Evie is proud to become a qualified gunner girl, part of the team operating the anti-aircraft batteries. When she meets RAF bomber pilot Matt Goodwin at a dance, it's not long before they fall in love. But then tragedy strikes. Matt is killed on a mission, leaving Evie bereft - and expecting a child.

1941, Poland. Though war rages around her and occupying German forces march steadily onward, 16-year-old Aleksandra has lived her life sheltered from danger. Until one day, she goes to school—never to return. Pushed roughly into a truck with other schoolgirls like her, a single ominous word tells her all she needs to know about her fate: Lebensborn.

After Pearl's home is destroyed in the Exeter Blitz, so too are her dreams of performing onstage. Finding work as a bus conductress instead, a chance encounter revives her hopes once more, and soon she is singing for the troops alongside new friend Ivy. When agent Gordon Gold approaches them, Ivy jumps to sign with him and sets off for the bright lights of London. But Pearl is wary of the charming man and decides to stay, watching her friend go with a heavy heart. A year later, while Pearl is struck mute by an illness, Ivy returns - and is quick to seize the chance to fill Pearl's place.

For Squadron Leader Alex Everton dogfights in his Spitfire have become his way of life. He knows every time he goes up in the air, he might not make it home again, but he’ll keep fighting anyway. The boys in blue seem to be winning the Battle of Britain, but during a raid over Dieppe the Luftwaffe gain the upper hand and Alex’s spitfire is shot down. Alex expects to either drown in the water beneath him or be incarcerated in a German prisoner of war camp. Instead, he becomes an evader, dodging the enemy as he tries to get back home to Blighty.

Jane’s parents think Eddie’s rebellious ways will lead their beloved daughter into trouble. They forbid her from seeing Eddie, telling Jane to get a proper job as a nurse. But Jane cannot stand the idea of life without the boy she loves and secretly continues to see Eddie. The day she finds out that Eddie has betrayed her, she is utterly distraught. Were her parents right all along?

1941. The Blitz rages over London. And even in Cornwall, the war is being fought. When Violet loses her sister in the Blitz, she must take her nieces to safety in Cornwall. On the coast, she meets carefree chorus girl Eva, who is also running from the dangers of London. But Porthcurno hides a secret military base, and soon Violet and Eva realise there’s a battle to fight in Cornwall, too. Together with local Hazel, who works on the base, they must come together to help the war effort. But will their friendship be enough to keep them safe? 

Kitty Wilson, who, after losing her father, rejects the unsuitable suitor her mother has chosen. Abandoned by her mother, Kitty faces the challenges of navigating life alone. Amidst the turmoil, Kitty finds solace and support from friends, securing a promotion at Harpers Emporium. However, tragedy looms, threatening to shatter her newfound stability. Sally and Ben Harper face their own challenges. A decision by Ben regarding the restaurant, made without consulting his partner Jack Burrows, jeopardizes not only their friendship but also the stability of Harpers itself.

21-year-old Alix is studying in Paris, far away from Sasha, her Serbian father and Leonora, her English mother, as World War Two breaks out. Finding hope even in troubled times, romance sparks with fellow student Raoul; but she soon learns young love can be tumultuous. So, when Steve, a handsome American pilot with Serbian ancestors, enters her life, she can't ignore her feelings toward him. And making a decision will be even harder as both men leave for war.

Cliffehaven, December 1946. From a hill above the town of Cliffehaven, a young woman makes a heart-breaking choice that will change the course of her life forever. Hours later, a baby is found in the Nativity crib of the local church. Who could have left him there, and why? The mystery preoccupies everyone in Cliffehaven - not least Peggy Reilly, who has enough to contend with at Beach View Boarding House without this extra secret to unravel.