As war rages, Barbara Sinclair is desperate to escape her unhappy home life. And with the threat of German bombs ever present, Barbara reluctantly agrees to marry John, her childhood friend, who is leaving to join the RAF. But an encounter with Alex Everton, a dashing Spitfire pilot, complicates matters for Barbara. With emotions running high, she begins to question whether she has made a terrible mistake. With the constant threat of death all around her, Barbara must try to find a way to deal with the complexities of her difficult home life and her emotional relationships, too.

When duty calls, East End girl Minnie Wolton isn’t afraid to do her bit. She’s excited to be joining the ATS girls and although she knows life at the barracks will be tough, she’s not one to quit! She just hopes the other girls are all good sorts. With her husband away fighting, Eileen Ruffel is glad of the break from her unhappy marriage. She just wants to make a success of the ATS girls and maybe find happiness too. Along with the other girls, Minnie and Eileen strike up a firm friendship. The training is hard, but they are determined to prove they are the best new recruits.

As the Nazi regime crumbles, Hanni hides from her father on the fringes of Berlin. She develops the photographs she took to record the brutality of the camp - the empty food bowls and hungry eyes - and vows to get justice for the innocent people she couldn’t help. But, on the day she plans to deliver these damning photographs to the Allies, Hanni comes face to face with her father again. Reiner Foss is now working with the British forces, his past safely hidden behind a new identity. He will go to deadly lengths to protect his secrets, but Hanni knows she can’t give up her fight. 

Autumn 1941, and as female conscription looms, three women arrive at the Bell Works Factory in Hucknall to work on the Spitfire production line. Lily soon realises her real dream is to pilot the planes; Edna longs to be reunited with her evacuee daughter, but soon discovers she has a much bigger problem; and Jeannie fears the worst after weeks of no news from her fiancé. Despite all the heartache, with a bit of festive cheer and 'can do' attitude, The Engine Girls will pull together and do all they can to ensure Britain's safe for the New Year.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

September 1939. London is in blackout; war has been declared, but Dulcie Treadwell can think only of her heartbreak over American broadcaster, Glenn Reeves, who didn’t say goodbye before leaving for Berlin. Posted to Bletchley Park, Dulcie must concentrate on helping the war effort by cracking the German Enigma codes. The hours are long and the conditions tough, with little recognition from above. Until she breaks her first code. Dulcie finally settles into something she was born to do. But just as she’s proving her worth, a shocking betrayal leads to Dulcie’s brutal dismissal.

Nazi-occupied Vienna, 1940. As bombs fall over Europe, Hannah Stern is a 27-year-old nurse who is just trying to survive. When she takes a position at Vienna’s esteemed psychiatric hospital, Am Steinhof, she thinks it will take her away from the danger. Her estranged half-brother has arranged the job for her on the condition that she must take his son, Willi, there too. He insists it’s a place where Willi will be safe. And Hannah soon forms a close bond with the sweet, fragile child in her care.

October 1940. When VAD nurse Evie narrowly escapes being killed in an air raid during the London Blitz, it propels her to make a life-changing decision to break free of her troubled and unhappy life. She escapes to the Norfolk countryside to start afresh, with a job at the newly opened Great Plumstead Hall hospital, and a wonderful new home at Rookery House. The community of Great Plumstead welcomes evacuees to the village—mothers and children bombed out of their London homes. Sisters Prue and Thea, along with members of The Mother's Day Club, help the new arrivals settle in.

Lady Joanna Harcourt and her daughter, Sarah, discover their life of luxury at Goodwill House is over. Because with Lord Harcourt away fighting, the Harcourt girls are facing financial ruin. Lady Joanna is terrified of losing their home, but for Sarah, this means her dreams of becoming a doctor are over. Headstrong Sarah isn’t one to quit! War or not, she’s determined to find a way to save her home and follow her dreams - whatever it takes.

Bristol, 1939. School leaver Maisie Miles suspects her father, a small-time crook, has an ulterior motive for insisting she gets a job at the W. D. & H. O. Wills tobacco factory but keeps it to herself. She's befriended by effervescent Phyllis Mason and kind-hearted Bridget Milligan who take pity on her and take Maisie under their wing. But beneath their happy-go-lucky exteriors, they all harbour dreams and worries about what the future holds.