It's 1938, and Cadi is chosen to be Rose Queen in the annual Rose Fete. She is thrilled to be treated like royalty for the day. But deep down she is desperate to leave the Welsh mining community where she grew up. When war is declared, Cadi and her best friend, Poppy, see a chance to escape. Cadi leaves behind her parents and local boy Aled, whom she is meant to marry, for Liverpool.

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.

Laura Collins is struggling to make a good life for her daughters, Tess and Tina. So when she is offered a good job, with accommodation, she is delighted. Life is very different for the Brewsters who live on Manor Farm in Herefordshire. Danny plans to marry one day so that he can take over the farm, whilst his cousin, Phil, longs to escape the world of farming as soon as possible. War changes all their plans.

Bristol, 1939. School leaver Maisie Miles suspects her father, a small-time crook, has an ulterior motive for insisting she gets a job at the W. D. & H. O. Wills tobacco factory but keeps it to herself. She's befriended by effervescent Phyllis Mason and kind-hearted Bridget Milligan who take pity on her and take Maisie under their wing. But beneath their happy-go-lucky exteriors, they all harbour dreams and worries about what the future holds. 

Lancashire, 1895. When her controlling stepfather suddenly dies, it seems that Elinor Pendleton finally has a chance of freedom. But her hopes are soon dashed when she learns that the thuggish Jason Stafford has inherited every penny, and is determined to have Elinor too. Forced to flee with her maid, Maude, Elinor finds shelter with Maude's distant cousin in the remote village on the shore of Jubilee Lake. But Walter Crossley has troubles of his own. Having lost his closest family in a tragic accident, he needs one of his grandsons to return from America to inherit his farm.

1850. When Opal Sharp finds herself and her younger siblings suddenly orphaned and destitute, she thinks things can get no worse. But soon three of them - including Opal - are struck down with the illness that took their father from them, and her brother Charlie is forced to make an impossible decision. Unable to afford a doctor, he knows the younger children will not survive. So, unbeknownst to Opal, Charlie takes their younger siblings to the workhouse, where he knows they will at least be fed and have a roof over their heads. Opal is heartbroken and struggles to forgive him. 

Hull, 1856. James Ripley and his wife Moira have always looked out for the poor of Hull. When, during one stormy night, there is a flood in a nearby cellar—a popular shelter for the homeless—James rushes to help. Among those rescued is a dark-haired little girl who speaks a language no one can understand. Some say that she came to the cellar with her mother, but no one knows where the mother is now. Concerned for the child's safety, James is unsure of what to do.

Autumn 1941, and as female conscription looms, three women arrive at the Bell Works Factory in Hucknall to work on the Spitfire production line. Lily soon realises her real dream is to pilot the planes; Edna longs to be reunited with her evacuee daughter, but soon discovers she has a much bigger problem; and Jeannie fears the worst after weeks of no news from her fiancé. Despite all the heartache, with a bit of festive cheer and 'can do' attitude, The Engine Girls will pull together and do all they can to ensure Britain's safe for the New Year.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

Abandoned by their parents and left to fight for themselves, headstrong Rosalind Carey has no choice but to take charge of her younger brothers and sister in Devonshire’s grand Rockwood Castle. But their once much-loved home is crumbling like the family that lives within it. Living hand to mouth and desperate to provide for the estate that depends on them, the Carey family are one debt away from ruin. Until the day comes when the dashing Piers Blanchard appears on their doorstep from Cornwall, claiming he is Rosalind’s distant cousin and that Rockwood Castle is his.

Cambridgeshire - March 1939. As the clouds of war begin to gather in Europe, the Talbot family of rural Blackberry Farm will be torn apart, just as so many families all over the world will be. Life will never be the same again. Whilst in London, the Salmons family will feel the pain of parting and loss. Brought together by war, the two families become intertwined and, as the outlook looks bleak, they must draw on each other’s strength to fight through the hard times.

London, 1939. When tragedy strikes, Libby Andrews is left homeless and destitute, fending for herself on the capital's most dangerous streets. Desperate times call for desperate measures, and Libby is on the cusp of making a decision that could jeopardise her future when a local boy saves her from ruin. The kindness of this stranger sets her on a different path, and Libby heads to Liverpool with a determined mind and hope in her heart. There, she reconnects with long lost family - but will she be able to uncover the truth that tore them apart all those years ago?

Scarborough, 1880. Young Jeannie spends her days watching her mother and the other harbour girls sitting at the water's edge - mending nets, gutting herring - and waiting for her friend Ethan Wharton to come in on his father's fishing smack. As she was growing up, Jeannie always expected to marry Ethan, who is loyal and dependable. But then she meets Harry - a stranger who has come to visit from Hull for the day - and she falls for him.

In a village in Wiltshire, just off a busy road, Larch Tree Lane winds slowly up a low hill. Among the residents is Maggie Hatherall, full of life despite her advancing years, who worries what she is going to do with her 200-year-old home. Lucia Grey has escaped a bad marriage but her ex is reluctant to let her start a new life free of him. And for architect Corin Drayton, an unexpected inheritance is the start of a complicated family wrangle. From the cottages and farms to the local shop, pub and old manor house, the residents' lives are set to intertwine and relationships bloom.

Lady Joanna Harcourt and her daughter, Sarah, discover their life of luxury at Goodwill House is over. Because with Lord Harcourt away fighting, the Harcourt girls are facing financial ruin. Lady Joanna is terrified of losing their home, but for Sarah, this means her dreams of becoming a doctor are over. Headstrong Sarah isn’t one to quit! War or not, she’s determined to find a way to save her home and follow her dreams - whatever it takes.