It's 1939, and the Brogan family of London's East End are ready to show Hitler what for. But things don't seem so rosy when rationing, evacuation and air raids start to put this larger-than-life family to the test. When a mysterious young man arrives at the Brogans' local parish church, he provides just the dazzling distraction they need - and, for eldest daughter Mattie, the promise of more than she'd ever wished for. But as the pair fall deeper in love, they are drawn into secret dangers, rife on the very London streets they call home.

When duty calls, East End girl Minnie Wolton isn’t afraid to do her bit. She’s excited to be joining the ATS girls and although she knows life at the barracks will be tough, she’s not one to quit! She just hopes the other girls are all good sorts. With her husband away fighting, Eileen Ruffel is glad of the break from her unhappy marriage. She just wants to make a success of the ATS girls and maybe find happiness too. Along with the other girls, Minnie and Eileen strike up a firm friendship. The training is hard, but they are determined to prove they are the best new recruits.

October 1940. When VAD nurse Evie narrowly escapes being killed in an air raid during the London Blitz, it propels her to make a life-changing decision to break free of her troubled and unhappy life. She escapes to the Norfolk countryside to start afresh, with a job at the newly opened Great Plumstead Hall hospital, and a wonderful new home at Rookery House. The community of Great Plumstead welcomes evacuees to the village—mothers and children bombed out of their London homes. Sisters Prue and Thea, along with members of The Mother's Day Club, help the new arrivals settle in.

After the hard work of being a maid at Wallington Manor in the lead-up to Christmas, Annie is very excited at the prospect of going home to Bethnal Green. She has missed her family, but the money she earns keeps them all afloat. Harriet is from the island of Guernsey and is visiting her aunt at Wallington Manor. When she has to leave for London Annie is asked to look after her. On the train journey a friendship blossoms. And when an accident happens their friendship is made even stronger. A friendship that would see both girls through pain, happiness, marriage and death.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

1949, County Durham. When Angie marries her sweetheart Quentin and moves into Cuthford Manor to begin their new life together, she feels like the luckiest woman in the world. But Quentin falls victim to a tragic accident and Angie's life is left devastated. Now, along with the prospect of rebuilding her life, she is faced with an impossible choice that will have far-reaching consequences for herself and those she loves most. Only with the help of her family, the community of Cuthford Manor and her old friends from the Shipyards, will Angie draw on the inner strength to find happiness again.

London, 1939. When tragedy strikes, Libby Andrews is left homeless and destitute, fending for herself on the capital's most dangerous streets. Desperate times call for desperate measures, and Libby is on the cusp of making a decision that could jeopardise her future when a local boy saves her from ruin. The kindness of this stranger sets her on a different path, and Libby heads to Liverpool with a determined mind and hope in her heart. There, she reconnects with long lost family - but will she be able to uncover the truth that tore them apart all those years ago?

Blackpool, 1893. Tilly has come a long way from the run-down tenements in which she grew up. She has a small but comfortable home, a loving, handsome husband, two beautiful little'uns - Babs and Beth - and she earns herself a little money weaving wicker baskets. Life is good. Until the day Tilly returns home to find a policeman standing on her doorstep. Her Arthur won't be coming home tonight - nor any night - having fallen to his death whilst working on Blackpool tower. Suddenly Tilly is her daughters' sole protector, and she's never felt more alone.

1940, Liverpool: Alice Turner and her best friend, Millie Markham, work for the war effort at Rootes munitions factory, making shell caps and Halifax bombers. Alice’s sweetheart, Terry, is home from the front for a brief period of leave. But the honeymoon is soon over. Terry must go back to the dangers of Nazi-occupied France, and the ever-present air raid sirens quickly bring Alice back down to earth. 

Dympna Byrne lives a simple life with her family on the Connemara coast. She adores her father Micheál, and does her best to help her family by working hard and expecting little. But beneath the surface there are hidden secrets. Dympna’s mother idolises her clever eldest son, yet her attitude to Dympna is puzzling. So when her family find themselves in desperate need of money, it is Dympna who crosses the water to Liverpool to seek a better life. 

Life for Elsie is difficult as she struggles to cope with her alcoholic mother. Caring for her siblings and working long hours at Swift's jam factory in London’s Bermondsey is exhausting. Thankfully her lifelong friendship with Dot helps to smooth over life’s rough edges. When Elsie and Dot meet Millie Swift, they are nervous to be in the presence of the bosses’ daughter. Over time, they are surprised to feel so drawn to her, but should two East End girls be socialising in such circles?

Rose McAllister is waiting for her husband, Steve, to come home. He is a seaman, often drunk and violent. Rose does her best to cope and sees that her daughters, Daisy and Petal, suffer as little as possible. Sensing that war is on the horizon, Steve tries to reform, but on his last night home, he pawns the girls' new dolls to go on a drinking binge. When war is declared, Rose finds a good job but agrees the children must be evacuated. War brings hardship and difficulty for the girls and Rose as they face the destruction of their home and cruelty at the hands of others....  

Nell Whitaker is 15 when war breaks out and her mother sends her to live with her Auntie Kath in a remote farmhouse on Anglesey. At first Nell hates it, but her friendship with Bryn, who works on the farm whilst waiting for his call-up papers, helps a lot. So when he leaves to join his ship, Nell is counting the days to his return. But her heart is broken when news reaches her that Bryn’s ship has been bombed and he has been drowned. Determined to bury her grief in hard work, Nell finds herself growing closer to Auntie Kath.

In February, 1922, at the western-most entrance to Victoria Station in Manchester, a massive plaque was unveiled. Beneath a vast tiled map showing the lines of the Lancashire and Yorkshire Railway network, a series of seven bronze panels recorded the names of the men of the Lancashire and Yorkshire Railway who gave their lives for King and Country in the Great War - a total of 1,460 names. In March, 1940, a group of women of varying ages and backgrounds stands in front of the memorial, ready to do its bit in this new World War.... 

September 1939. London is in blackout; war has been declared, but Dulcie Treadwell can think only of her heartbreak over American broadcaster, Glenn Reeves, who didn’t say goodbye before leaving for Berlin. Posted to Bletchley Park, Dulcie must concentrate on helping the war effort by cracking the German Enigma codes. The hours are long and the conditions tough, with little recognition from above. Until she breaks her first code. Dulcie finally settles into something she was born to do. But just as she’s proving her worth, a shocking betrayal leads to Dulcie’s brutal dismissal.