In 2015, a race of alien Others conquered Earth. They enslaved humanity not by force, but through an aggressive mind control that turned people into contented, unquestioning robots.
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Althea and Lucas barely escaped the Others' clutches in the autumn, and were separated in the process. Alone and on the run from the cruel alien race determined to exterminate her, Althea struggles to adapt and survive in a world she never imagined. When a boy named Pax appears out of nowhere, he quickly recognizes Althea for what she is - a human/Other hybrid just like him. Althea begs him to help her find Lucas, but Pax refuses, intent on following his own mysterious agenda.
Althea assumes now that she, Pax, and Lucas are reunited in spring that the next steps are obvious - locate Deshi and prepare to take down the Others once and for all. But she doesn't expect the subtle changes in Lucas. After being left alone last season with only his Element father for company, Lucas has started to question whether their rightful place isn't with the strange alien race as opposed to humanity.
A final battle for the survival of Earth is coming. Between the alien Others and the destruction of humanity stand four Dissidents. When the Prime Other banishes them to the Harvest Site to live the remainder of their time on Earth as slaves, the Dissidents use the opportunity to learn more about the substance that keeps the Others alive...and how they might use it to their advantage.