Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

As war rages, Barbara Sinclair is desperate to escape her unhappy home life. And with the threat of German bombs ever present, Barbara reluctantly agrees to marry John, her childhood friend, who is leaving to join the RAF. But an encounter with Alex Everton, a dashing Spitfire pilot, complicates matters for Barbara. With emotions running high, she begins to question whether she has made a terrible mistake. With the constant threat of death all around her, Barbara must try to find a way to deal with the complexities of her difficult home life and her emotional relationships, too.

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.

London, 1912. Identical twins Lester and Lillia Holdsworth are destined for the stage. Lester is a brilliant pianist, Lillia a magnificent opera singer. But their father has other ideas. Lester is sent to a military academy while Lillia must marry Lord Dalton - a self-righteous, pompous friend of her father's looking for a young wife to give him an heir. Yet their plans to defy their father's wishes are put on hold when war breaks out in 1914.

1940. In a world newly burning with war, and in spite of her American family’s wishes, Virginia d’Albert-Lake decides to stay in occupied France with her French husband. She’s sure that if they keep their heads down, they’ll survive. But is surviving enough? Nineteen-year-old Violette Szabo has seen the Nazis’ evil up close and is desperate to fight them. But when she meets the man who’ll change her life only for tragedy to strike, Violette’s adrift. 

As if everyday life in London's slums wasn't hard enough for Jane Roberts and her three children, the day that ne'er-do-well husband Bert walks out on his family as rent day draws near is a particularly dark time. In the wake of his desertion, the family pulls together to find their way forward, and Jane takes the steps she needs to get a divorce and free them all from Bert in the future. With help from a neighbour who is not all she seems, a lucky break and fresh opportunities, the road ahead starts to look more promising. 

When Stefania Carmichael steps into the mysterious world of Bletchley Park, she immediately finds herself signing the Official Secrets Act. In whispers, she is told that she’s been recruited because of her talent for languages. Before the war ripped Europe apart, Stefania was living in Rome, charming everyone she met, and engaged to the man of her dreams, handsome and brave Matteo. And now everything has changed. With secrets swirling around the building, Stefania finds it hard to know who to trust, until she forms a close bond with two other new recruits.

The war has ended; the men are returning home to their loved ones, but for some, things have changed. Charlotte Hennessey-White's husband, David, is no longer the gentle loving man he once was and Charlotte, so independent during the war, is devastated. Edna Burbage's strong fiancée, Colin, has suffered appalling physical injuries. He won’t hold her to her promise of marriage, but she insists her feelings are unchanged. But is that true? And Polly Chandler's sweetheart, Gavin, who’d she’d planned her whole future around, hasn’t come home at all.

Maureen Jackson knew life as a trainee nurse wouldn't be easy, but she didn't expect her hospital to be badly bombed on her first shift. Plus Maureen still has her family and friends in Mulberry Lane to keep her busy - she's needed as much there as she is by her patients. Running the pub on the corner of Mulberry Lane, Peggy Ashley is used to taking in all sorts of waifs and strays. But the arrival of a dashing American captain has got tongues wagging about Mulberry Lane's favourite landlady....

Cambridgeshire - March 1939. As the clouds of war begin to gather in Europe, the Talbot family of rural Blackberry Farm will be torn apart, just as so many families all over the world will be. Life will never be the same again. Whilst in London, the Salmons family will feel the pain of parting and loss. Brought together by war, the two families become intertwined and, as the outlook looks bleak, they must draw on each other’s strength to fight through the hard times.