Germany, 1937. Fear and betrayal stalk the streets. People disappear. Persecution of the Jews has become a national pastime. When Ruth Friedman's husband is arrested by the SS, she is left to fend for herself and her four children. She alone stands as their shield against the Nazis. But where can she go? Where will her family be safe? Ruth must overcome the indifference, hatred and cruelty that surrounds her as she and her family race to escape the advancing Nazi army's final solution.

It is two years before the outbreak of World War 1, and the Oakley family are typical of their time. Andrew has been clerk at a City solicitors for twenty years. Alice, his wife, keeps the home fires burning. Their sons plan on becoming skilled apprentices. And Mabel, their daughter, says she would rather die than go into domestic service like her cousin. She is going to go to train as a secretary. Andrew Oakley little imagines, as he sets off for work one February morning, that their lives are about to change in the most dramatic and tragic fashion.

Dartmouth Circle is quiet and exclusive - just the way its residents want it to stay, especially arch busybody Sheila Colby. When she hears that the house next door to her has been bought for a student and four friends to rent, she is incandescent. Nothing in this outwardly serene little cul-de-sac is quite as it seems - will the arrival of Madeleine 'Mad' Richmond and her friends bring seething tensions and dark secrets bubbling to the surface?

Bristol 1939. At forty years of age, Mary Anne Randall still turns heads. With an abusive husband spending most of his wages on beer, she has resigned herself to be the sole breadwinner to protect her family and keep the wolves from the door. In order to make ends meet, Mary Anne runs a pawnbroking business from the wash house at the end of the garden. Following the outbreak of war an opportunity presents itself to escape her loveless marriage and find true love. Will she take that chance, or carry on living just for her children and not for herself?

Wakefield 1870. On the banks of the River Calder, Meg Taylor struggles to care for her younger siblings and sick mother whilst her father is away working as a boatman on the canals. The slums where they live are rife with disease, and Meg longs for a different life, away from the grinding poverty, but she'll never leave her family. But with the canals slowly dying as trade moves to the railways, and with Meg’s father stubbornly holding onto the past, Meg fears her family’s future is going to get even bleaker. If only there was a way she could save them.

England, 1943: Home is no longer safe for eight-year-old twins Molly and Jacob. Night after night, wailing bombs and screeching planes skim the rooftops overhead. They cradle each other, shivering in terror, not knowing if they'll live to see dawn. Their mother, Martha, has no choice but to evacuate them to the safety of the countryside. At the train station, Martha bites back tears as she says good-bye to her precious children. Knowing she might never see them again, she gives Jacob a letter, pressing the envelope into his hands and telling him to only read it if they are in danger.

England, 1941: Life has always been cruel to Ethel. Raised in an orphanage by a merciless mistress, she never knew the meaning of love. And now she has fallen for the one man who is forbidden to her - Karl, a German held captive in the local prisoner-of-war camp. He might be the enemy, but he has her heart. When Ethel discovers she’s pregnant, she is labelled a traitor and shunned by all. The love of her life is torn away from her, but she vows to protect their precious daughter. But the unthinkable happens....

1940, Liverpool: Alice Turner and her best friend, Millie Markham, work for the war effort at Rootes munitions factory, making shell caps and Halifax bombers. Alice’s sweetheart, Terry, is home from the front for a brief period of leave. But the honeymoon is soon over. Terry must go back to the dangers of Nazi-occupied France, and the ever-present air raid sirens quickly bring Alice back down to earth. 

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.

Virginia, 1933: Lauren Greenwood was born to look pretty and land a rich husband. For her and her high-society family, it’s easy to forget that America is in the devastating grip of the Great Depression, where children run wild in the streets, endless queues for soup kitchens line sidewalks, and desperation hangs in the air. But when a young servant comes to Lauren, pregnant and begging for help, her world crumbles. Someone she loves is responsible, and when she finds out the shocking truth she must decide: Protect the Greenwood name or be cast out? 

Liverpool, 1939. Bella Rogers thought her life was just starting; she can’t wait to get married to her childhood sweetheart, Bobby. But when war is declared and Bobby is called up, his letters stop, leaving Bella heartbroken and alone. Then her family is torn apart by tragedy when her angelic 5-year-old sister dies from a terrible illness. They can barely afford a funeral, but the church is packed with the entire community, devastated for their loss. Grieving for her sister and lost love, Bella finds a job at the Bryant and May factory, making matches.

Bristol, 1939. School leaver Maisie Miles suspects her father, a small-time crook, has an ulterior motive for insisting she gets a job at the W. D. & H. O. Wills tobacco factory but keeps it to herself. She's befriended by effervescent Phyllis Mason and kind-hearted Bridget Milligan who take pity on her and take Maisie under their wing. But beneath their happy-go-lucky exteriors, they all harbour dreams and worries about what the future holds. 

The war has ended; the men are returning home to their loved ones, but for some, things have changed. Charlotte Hennessey-White's husband, David, is no longer the gentle loving man he once was and Charlotte, so independent during the war, is devastated. Edna Burbage's strong fiancée, Colin, has suffered appalling physical injuries. He won’t hold her to her promise of marriage, but she insists her feelings are unchanged. But is that true? And Polly Chandler's sweetheart, Gavin, who’d she’d planned her whole future around, hasn’t come home at all.

In a village in Wiltshire, just off a busy road, Larch Tree Lane winds slowly up a low hill. Among the residents is Maggie Hatherall, full of life despite her advancing years, who worries what she is going to do with her 200-year-old home. Lucia Grey has escaped a bad marriage but her ex is reluctant to let her start a new life free of him. And for architect Corin Drayton, an unexpected inheritance is the start of a complicated family wrangle. From the cottages and farms to the local shop, pub and old manor house, the residents' lives are set to intertwine and relationships bloom.

Taken in by the firm but kind-hearted cook, the young friends can hardly believe their luck. But behind Bracken House’s impressive façade lies a household steeped in troubles and mystery, with residents above and below stairs battling their own demons and dark secrets. Not everyone is happy about the new arrivals, and soon the orphans’ safety is in danger. If they want to stay in the first home any of them have known for years, they must unravel the past and bring hope to the future. Will they succeed? Or will they come to regret ever leaving the mean slum streets they once called home?