It's 1939, and the Brogan family of London's East End are ready to show Hitler what for. But things don't seem so rosy when rationing, evacuation and air raids start to put this larger-than-life family to the test. When a mysterious young man arrives at the Brogans' local parish church, he provides just the dazzling distraction they need - and, for eldest daughter Mattie, the promise of more than she'd ever wished for. But as the pair fall deeper in love, they are drawn into secret dangers, rife on the very London streets they call home.

When duty calls, East End girl Minnie Wolton isn’t afraid to do her bit. She’s excited to be joining the ATS girls and although she knows life at the barracks will be tough, she’s not one to quit! She just hopes the other girls are all good sorts. With her husband away fighting, Eileen Ruffel is glad of the break from her unhappy marriage. She just wants to make a success of the ATS girls and maybe find happiness too. Along with the other girls, Minnie and Eileen strike up a firm friendship. The training is hard, but they are determined to prove they are the best new recruits.

October 1940. When VAD nurse Evie narrowly escapes being killed in an air raid during the London Blitz, it propels her to make a life-changing decision to break free of her troubled and unhappy life. She escapes to the Norfolk countryside to start afresh, with a job at the newly opened Great Plumstead Hall hospital, and a wonderful new home at Rookery House. The community of Great Plumstead welcomes evacuees to the village—mothers and children bombed out of their London homes. Sisters Prue and Thea, along with members of The Mother's Day Club, help the new arrivals settle in.

After the hard work of being a maid at Wallington Manor in the lead-up to Christmas, Annie is very excited at the prospect of going home to Bethnal Green. She has missed her family, but the money she earns keeps them all afloat. Harriet is from the island of Guernsey and is visiting her aunt at Wallington Manor. When she has to leave for London Annie is asked to look after her. On the train journey a friendship blossoms. And when an accident happens their friendship is made even stronger. A friendship that would see both girls through pain, happiness, marriage and death.

As war rages, Barbara Sinclair is desperate to escape her unhappy home life. And with the threat of German bombs ever present, Barbara reluctantly agrees to marry John, her childhood friend, who is leaving to join the RAF. But an encounter with Alex Everton, a dashing Spitfire pilot, complicates matters for Barbara. With emotions running high, she begins to question whether she has made a terrible mistake. With the constant threat of death all around her, Barbara must try to find a way to deal with the complexities of her difficult home life and her emotional relationships, too.

London, 1939. When tragedy strikes, Libby Andrews is left homeless and destitute, fending for herself on the capital's most dangerous streets. Desperate times call for desperate measures, and Libby is on the cusp of making a decision that could jeopardise her future when a local boy saves her from ruin. The kindness of this stranger sets her on a different path, and Libby heads to Liverpool with a determined mind and hope in her heart. There, she reconnects with long lost family - but will she be able to uncover the truth that tore them apart all those years ago?

Blackpool, 1893. Tilly has come a long way from the run-down tenements in which she grew up. She has a small but comfortable home, a loving, handsome husband, two beautiful little'uns - Babs and Beth - and she earns herself a little money weaving wicker baskets. Life is good. Until the day Tilly returns home to find a policeman standing on her doorstep. Her Arthur won't be coming home tonight - nor any night - having fallen to his death whilst working on Blackpool tower. Suddenly Tilly is her daughters' sole protector, and she's never felt more alone.

Alice is nursing an injured hand and a broken heart when she moves to the village of Churchwood at the start of WWII. She is desperate to be independent but worries that her injuries will make that impossible. Kate lives with her family on Brimbles Farm, where her father and brothers treat her no better than a servant. With no mother or sisters, and shunned by the locals, Kate longs for a friend of her own. Naomi is looked up to for owning the best house in the village. But privately, she carries the hurts of childlessness.

Dympna Byrne lives a simple life with her family on the Connemara coast. She adores her father Micheál, and does her best to help her family by working hard and expecting little. But beneath the surface there are hidden secrets. Dympna’s mother idolises her clever eldest son, yet her attitude to Dympna is puzzling. So when her family find themselves in desperate need of money, it is Dympna who crosses the water to Liverpool to seek a better life. 

Life for Elsie is difficult as she struggles to cope with her alcoholic mother. Caring for her siblings and working long hours at Swift's jam factory in London’s Bermondsey is exhausting. Thankfully her lifelong friendship with Dot helps to smooth over life’s rough edges. When Elsie and Dot meet Millie Swift, they are nervous to be in the presence of the bosses’ daughter. Over time, they are surprised to feel so drawn to her, but should two East End girls be socialising in such circles?

Liverpool, 1942. Though they may be posted miles apart, Cadi and Jez's love for one another is stronger than ever—until Cadi receives a heart-breaking call from Jez's base. Cadi races to be with her beau and at his bedside, they make each other a promise before they say their goodbyes. In her darkest hour, Cadi is comforted by her friends and with their support and guidance she begins to see a light at the end of the tunnel. But Cadi will face more than one unexpected turn along the way.

Rose McAllister is waiting for her husband, Steve, to come home. He is a seaman, often drunk and violent. Rose does her best to cope and sees that her daughters, Daisy and Petal, suffer as little as possible. Sensing that war is on the horizon, Steve tries to reform, but on his last night home, he pawns the girls' new dolls to go on a drinking binge. When war is declared, Rose finds a good job but agrees the children must be evacuated. War brings hardship and difficulty for the girls and Rose as they face the destruction of their home and cruelty at the hands of others....  

In February, 1922, at the western-most entrance to Victoria Station in Manchester, a massive plaque was unveiled. Beneath a vast tiled map showing the lines of the Lancashire and Yorkshire Railway network, a series of seven bronze panels recorded the names of the men of the Lancashire and Yorkshire Railway who gave their lives for King and Country in the Great War - a total of 1,460 names. In March, 1940, a group of women of varying ages and backgrounds stands in front of the memorial, ready to do its bit in this new World War.... 

Nell Whitaker is 15 when war breaks out and her mother sends her to live with her Auntie Kath in a remote farmhouse on Anglesey. At first Nell hates it, but her friendship with Bryn, who works on the farm whilst waiting for his call-up papers, helps a lot. So when he leaves to join his ship, Nell is counting the days to his return. But her heart is broken when news reaches her that Bryn’s ship has been bombed and he has been drowned. Determined to bury her grief in hard work, Nell finds herself growing closer to Auntie Kath.

August 1940: As the Luftwaffe swarm over Liverpool, Shane Quinn decides to move his family back to the safety of Ireland. But his only child, the beautiful Dana, would rather stay and serve her country than flee to a foreign land. Determined to make it on her own, she joins the WAAF with newfound pals Patty and Lucy. There’s plenty of excitement to be had on a RAF station, even a chance or two at love.