Lady Joanna Harcourt and her daughter, Sarah, discover their life of luxury at Goodwill House is over. Because with Lord Harcourt away fighting, the Harcourt girls are facing financial ruin. Lady Joanna is terrified of losing their home, but for Sarah, this means her dreams of becoming a doctor are over. Headstrong Sarah isn’t one to quit! War or not, she’s determined to find a way to save her home and follow her dreams - whatever it takes.

Jane Hadley has nothing to lose when she runs away to join the Women's Auxiliary Air Force. Whatever faces her in war-torn London can't be any worse than staying at home with her abusive father. The city is nothing like she could have imagined, but she's soon on the move, travelling from base to base for her top-secret training. Making plenty of friends along the way, it doesn't take long for Jane to embrace her growing confidence - especially under the attentive eye of dashing Officer Oscar Stanton.

As war rages, Barbara Sinclair is desperate to escape her unhappy home life. And with the threat of German bombs ever present, Barbara reluctantly agrees to marry John, her childhood friend, who is leaving to join the RAF. But an encounter with Alex Everton, a dashing Spitfire pilot, complicates matters for Barbara. With emotions running high, she begins to question whether she has made a terrible mistake. With the constant threat of death all around her, Barbara must try to find a way to deal with the complexities of her difficult home life and her emotional relationships, too.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

The war has ended; the men are returning home to their loved ones, but for some, things have changed. Charlotte Hennessey-White's husband, David, is no longer the gentle loving man he once was and Charlotte, so independent during the war, is devastated. Edna Burbage's strong fiancée, Colin, has suffered appalling physical injuries. He won’t hold her to her promise of marriage, but she insists her feelings are unchanged. But is that true? And Polly Chandler's sweetheart, Gavin, who’d she’d planned her whole future around, hasn’t come home at all.

Bristol - 1953. It’s Coronation Year. A new beginning in the aftermath of war, but there are still battles to be fought and secrets to be kept. Charlotte Hennessey-White copes with the shortcomings of her marriage and throws herself into helping refugees unwelcome by some and exploited by others. Edna Burbage has three beautiful children and considers herself lucky until the advent of a deadly twentieth century disease makes her think otherwise. Polly Chandler still hopes for a better life, but there are too many obstacles standing in her way.

Bristol 1939. At forty years of age, Mary Anne Randall still turns heads. With an abusive husband spending most of his wages on beer, she has resigned herself to be the sole breadwinner to protect her family and keep the wolves from the door. In order to make ends meet, Mary Anne runs a pawnbroking business from the wash house at the end of the garden. Following the outbreak of war an opportunity presents itself to escape her loveless marriage and find true love. Will she take that chance, or carry on living just for her children and not for herself?

September 1939. London is in blackout; war has been declared, but Dulcie Treadwell can think only of her heartbreak over American broadcaster, Glenn Reeves, who didn’t say goodbye before leaving for Berlin. Posted to Bletchley Park, Dulcie must concentrate on helping the war effort by cracking the German Enigma codes. The hours are long and the conditions tough, with little recognition from above. Until she breaks her first code. Dulcie finally settles into something she was born to do. But just as she’s proving her worth, a shocking betrayal leads to Dulcie’s brutal dismissal.

London, 1941. The Blitz. When a Royal Navy memo arrives at head office, requesting female recruits to sign up as motorcycle dispatch riders, delivering highly classified orders across the country, three women jump at the chance to sign up for the most dangerous jobs in London.

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.

At the heart of Dover's Market Square lies the Castle's Café. A bustling, homely place run by the formidable Nellie Castle and her six children. Since the scandalous birth of her beloved son 10 years before, Marianne, Nellie's eldest daughter, has preferred to stay in the kitchen, hidden away from the town gossips. Overcome with shame, Marianne has never revealed the name of Donny's father - not even to her own mother. But with World War II just around the corner, Marianne's past catches up with her. And suddenly the lives of the Castle family become a lot more complicated.

Cambridgeshire - March 1939. As the clouds of war begin to gather in Europe, the Talbot family of rural Blackberry Farm will be torn apart, just as so many families all over the world will be. Life will never be the same again. Whilst in London, the Salmons family will feel the pain of parting and loss. Brought together by war, the two families become intertwined and, as the outlook looks bleak, they must draw on each other’s strength to fight through the hard times.

Hull, 1940. Widowed Edie Copeland has just arrived on Exchange Street, heavily pregnant and with heavy suitcase in tow. She left York after tragedy struck to make a fresh start, but she's a stranger to this street, and her new landlord doesn't hesitate to remind her of this, widow or not. Luckily, the neighbours are a little more welcoming and Edie is soon made to feel at home by the Maguires and the Scuttles. As air raids sound and the war feels closer than ever, the community has to stick together. But Edie is hiding something, and she doesn't know how much longer her secret is safe.

In February, 1922, at the western-most entrance to Victoria Station in Manchester, a massive plaque was unveiled. Beneath a vast tiled map showing the lines of the Lancashire and Yorkshire Railway network, a series of seven bronze panels recorded the names of the men of the Lancashire and Yorkshire Railway who gave their lives for King and Country in the Great War - a total of 1,460 names. In March, 1940, a group of women of varying ages and backgrounds stands in front of the memorial, ready to do its bit in this new World War.... 

1941. The Blitz rages over London. And even in Cornwall, the war is being fought. When Violet loses her sister in the Blitz, she must take her nieces to safety in Cornwall. On the coast, she meets carefree chorus girl Eva, who is also running from the dangers of London. But Porthcurno hides a secret military base, and soon Violet and Eva realise there’s a battle to fight in Cornwall, too. Together with local Hazel, who works on the base, they must come together to help the war effort. But will their friendship be enough to keep them safe?