September 1939. London is in blackout; war has been declared, but Dulcie Treadwell can think only of her heartbreak over American broadcaster, Glenn Reeves, who didn’t say goodbye before leaving for Berlin. Posted to Bletchley Park, Dulcie must concentrate on helping the war effort by cracking the German Enigma codes. The hours are long and the conditions tough, with little recognition from above. Until she breaks her first code. Dulcie finally settles into something she was born to do. But just as she’s proving her worth, a shocking betrayal leads to Dulcie’s brutal dismissal.

In February, 1922, at the western-most entrance to Victoria Station in Manchester, a massive plaque was unveiled. Beneath a vast tiled map showing the lines of the Lancashire and Yorkshire Railway network, a series of seven bronze panels recorded the names of the men of the Lancashire and Yorkshire Railway who gave their lives for King and Country in the Great War - a total of 1,460 names. In March, 1940, a group of women of varying ages and backgrounds stands in front of the memorial, ready to do its bit in this new World War.... 

When Stefania Carmichael steps into the mysterious world of Bletchley Park, she immediately finds herself signing the Official Secrets Act. In whispers, she is told that she’s been recruited because of her talent for languages. Before the war ripped Europe apart, Stefania was living in Rome, charming everyone she met, and engaged to the man of her dreams, handsome and brave Matteo. And now everything has changed. With secrets swirling around the building, Stefania finds it hard to know who to trust, until she forms a close bond with two other new recruits.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

When duty calls, East End girl Minnie Wolton isn’t afraid to do her bit. She’s excited to be joining the ATS girls and although she knows life at the barracks will be tough, she’s not one to quit! She just hopes the other girls are all good sorts. With her husband away fighting, Eileen Ruffel is glad of the break from her unhappy marriage. She just wants to make a success of the ATS girls and maybe find happiness too. Along with the other girls, Minnie and Eileen strike up a firm friendship. The training is hard, but they are determined to prove they are the best new recruits.

It is 1940s coastal Lincolnshire, and Carolyn Holmes is keen to do what she can for the war effort. Raised on the family farm, she battles with her mother to further her education, although nothing is too good for her brother. Phyllis Carter, a bitter widow from the Great War, lives close by with her son, who works on the farm. When Peter decides to volunteer, a distraught Phyllis blames Carolyn, who leaves to join the ATS where she meets Beryl. Carolyn and Beryl are posted to Beaumanor Hall in Leicestershire as ‘listeners’, the most difficult of signals intelligence gathering.

Jane’s parents think Eddie’s rebellious ways will lead their beloved daughter into trouble. They forbid her from seeing Eddie, telling Jane to get a proper job as a nurse. But Jane cannot stand the idea of life without the boy she loves and secretly continues to see Eddie. The day she finds out that Eddie has betrayed her, she is utterly distraught. Were her parents right all along?

Berlin, 1941: Margarete Rosenbaum is working as a housemaid for a senior Nazi officer when his house is bombed, leaving her the only survivor. But when she’s mistaken for his daughter in the aftermath of the blast, Margarete knows she can make a bid for freedom.... Issued with temporary papers - and with the freedom of not being seen as Jewish - a few hours are all she needs to escape to relative safety. That is, until her former employer’s son, SS officer Wilhelm Huber, tracks her down.

Bristol, 1939. School leaver Maisie Miles suspects her father, a small-time crook, has an ulterior motive for insisting she gets a job at the W. D. & H. O. Wills tobacco factory but keeps it to herself. She's befriended by effervescent Phyllis Mason and kind-hearted Bridget Milligan who take pity on her and take Maisie under their wing. But beneath their happy-go-lucky exteriors, they all harbour dreams and worries about what the future holds. 

October 1940. When VAD nurse Evie narrowly escapes being killed in an air raid during the London Blitz, it propels her to make a life-changing decision to break free of her troubled and unhappy life. She escapes to the Norfolk countryside to start afresh, with a job at the newly opened Great Plumstead Hall hospital, and a wonderful new home at Rookery House. The community of Great Plumstead welcomes evacuees to the village—mothers and children bombed out of their London homes. Sisters Prue and Thea, along with members of The Mother's Day Club, help the new arrivals settle in.

There are some things which even the closest friendship cannot survive.... Neighbours Edie Kelsey and Lil Horton have been friends for more than twenty years, sharing the joys and sorrows of tough lives as the wives of fishermen in Grimsby. So it was no surprise that their children were close and that Edie's son, Frank, and Lil's daughter, Irene, would fall in love and marry at a young age. But the declaration of war in 1939 changes everything.

London 1941. Sirens warn of danger as German bomber planes approach the city. Ever cautious, Kathy and her mother head immediately for the nearest public air raid shelter, and when the building is destroyed in a direct hit, it is a miracle Kathy survives. Finding herself alone, Kathy is left with nothing but a burning fury and a determination to join the war effort. 

Ana Kaminski is pushed through the iron gates of Auschwitz beside her frightened young friend Ester Pasternak. As they reach the front of the line, Ana steps forward and quietly declares herself a midwife--and Ester her assistant. Their arms are tattooed and they’re ordered to the maternity hut. Holding an innocent newborn baby, Ana knows the fate of so many are in her hands and vows to do everything she can to save them.

It is 1937, and Alice, Edith and Marg continue to face hardships every day, growing up on one of the poorest streets in Blackpool. Penniless, their friendship has helped them survive this far, but it'll take more than that to see them through the dark days ahead. Alice is coping with a violent father. Marg is left reeling after a dark secret about her birth comes to light and threatens to destroy the life she knows, and Edith is fighting to protect her alcoholic mother from the shame of their neighbours and keep her brother on the straight and narrow. 

Robinswood, Co. Waterford, 1939. The once grand house is home to two very different families. Despite delusions of grandeur, Lord and Lady Kenefick and their adult children live a life of decayed opulence as the money needed to keep such a large house and grounds ever dwindles. Meanwhile, the Murphy family, Dermot, Isabella and their three almost grown-up girls, live and work on the estate and do their best to keep everything running smoothly.