When duty calls, East End girl Minnie Wolton isn’t afraid to do her bit. She’s excited to be joining the ATS girls and although she knows life at the barracks will be tough, she’s not one to quit! She just hopes the other girls are all good sorts. With her husband away fighting, Eileen Ruffel is glad of the break from her unhappy marriage. She just wants to make a success of the ATS girls and maybe find happiness too. Along with the other girls, Minnie and Eileen strike up a firm friendship. The training is hard, but they are determined to prove they are the best new recruits.

Kate Bratton dreams of more. It’s 1914, and her life is mapped out ahead of her: continue working as a maid in the beautiful Woodicombe House, settle down with Luke the gardener and, of course, start a family. Desperate to run away in search of adventure, Kate’s plans are curtailed by the arrival of the Russell family at Woodicombe House. Tasked with becoming a lady’s maid for their daughter, Naomi, Kate gets a glimpse of the other side of life. Will she return to the safety of her life before, or will the handsome Ned Russell turn her head?

1914. Evie Munro is a Scottish fisher girl, working the herring season from Wick to Great Yarmouth. For Evie, every day is the same - gutting fish at the docks, shoulder-to-shoulder with her friends, followed by fresh bread, a warm whiskey toddy and an early night. But when Germany declares war, everything changes. As her village begins to empty of young men, Evie's life is marked by a heartbreaking tragedy. Her happiness destroyed, she vows to join the war effort as an army nurse, caring for wounded soldiers on the imposing hospital ship Britannic.

Left homeless after the Exeter Blitz, May must find a job if she's to put a roof over her head and help support her two younger sisters. Taking a job as housekeeper to farmer George, May soon ends up getting stuck into more than just cleaning and cooking. The Ministry of Agriculture will close Fair Maids Farm if it doesn't meet their produce targets, but George refuses to heed their warnings. With only two reluctant Land Girls to help, May receives unexpected guidance from Dan, a neighbouring farmer, whose kindness gets tongues wagging in the village.

As war rages, Barbara Sinclair is desperate to escape her unhappy home life. And with the threat of German bombs ever present, Barbara reluctantly agrees to marry John, her childhood friend, who is leaving to join the RAF. But an encounter with Alex Everton, a dashing Spitfire pilot, complicates matters for Barbara. With emotions running high, she begins to question whether she has made a terrible mistake. With the constant threat of death all around her, Barbara must try to find a way to deal with the complexities of her difficult home life and her emotional relationships, too.

1942. Evie Sinclair has enlisted in the ATS to serve her country. Her mother disapproves, wanting her to stay and help run the household, but her father supports her bid for independence. Though conditions are austere and the training is intense, Evie is proud to become a qualified gunner girl, part of the team operating the anti-aircraft batteries. When she meets RAF bomber pilot Matt Goodwin at a dance, it's not long before they fall in love. But then tragedy strikes. Matt is killed on a mission, leaving Evie bereft - and expecting a child.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

When the love of Tasha’s life, Luca, joins the air force to fight against the evil Nazi invaders, she knows she has to follow her heart—and him—into battle. Headstrong, impulsive and a daredevil, she’s the perfect recruit. Tasha’s sensible older sister, Nadia, plans only to stop Tasha’s madness and bring her home. But a chance encounter puts her in a plane, soaring above the clouds, and she also finds her calling.

East London, 1940. At the outbreak of war, hopeless romantic Prue Carmichael and her sister must leave their rural parish behind when their father is posted to a church in Stepney. To Prue, the blitz-ravaged streets of London seem an unlikely place for love to bloom – until a chance encounter throws rakish engineer Jack Quinn into her path. But as their connection deepens, his troubled past begins to emerge, and Prue realises Jack has secrets to hide.

Cornwall, England, 1941. Gazing at the coiled barbed wire and danger signs on this once-beautiful beach, Maddie suddenly hears a frightened little cry. Her heart pounding, she races down the dunes, but knows she will never reach the child in time. Maddie Brady is overwhelmed with relief to see a brave soul ahead of her rescue a little boy trapped among the landmines on the beach. But thanking the man, Maddie is shocked to hear a German accent. Why would an escaped German prisoner of war rescue an English boy? 

Following a humiliating experience involving the man she thought she’d marry, Iris Tredwick signs up to the Wrens in order to escape and - hopefully - find ‘the right sort’ of man to please her mother. After a bumpy start, Iris manages to befriend outspoken Mary and dreamer Sally as they are sent to their first posting - in Orkney. There she meets engineer Rob, whose flirtatious nature both charms and confounds straight-laced Iris. Much more appropriate for her is local doctor Stewart, if only she felt the same spark for him as she does for Rob.

England, 1940. Opening the box with trembling hands, she couldn’t believe it–a tiny baby lay inside. She gently lifted the newborn as it started to cry. Cradling the little one to her chest, she searched the darkness for any sign of whoever left it here. And as she rocked the child, something fluttered to the ground...a ration book.

Essex 1941. Unhappy with life at home with her privileged family, Hannah Austen-Bagshaw runs away to join the Land Girls Army, carrying with her a dark and terrible secret. Life on the farm is tough, but when Hannah meets handsome working-class flight lieutenant Jack, they form an immediate friendship. But unbeknownst to Jack, Hannah is sheltering a young German pilot who has crash landed in a nearby field. Torn between her duty and her humanity, Hannah knows she risks being arrested as a traitor, but she cannot see the young pilot suffer.

1944. With deadly doodlebugs terrorising London, Bessie Rushbrook agrees to give shelter to eight-year-old evacuee Marigold in her home. But, the little girl is no stranger, and Bessie must honour the promise she gave to Marigold's mother and not reveal her connection to the child. Marigold's arrival stirs up Bessie's memories from the past when the world was at war the first time. She is forced to face her actions from those days and question the secret that she's long kept hidden. When Grace is injured in London, Bessie makes the heart-wrenching decision to confess her secret.

Lady Joanna Harcourt and her daughter, Sarah, discover their life of luxury at Goodwill House is over. Because with Lord Harcourt away fighting, the Harcourt girls are facing financial ruin. Lady Joanna is terrified of losing their home, but for Sarah, this means her dreams of becoming a doctor are over. Headstrong Sarah isn’t one to quit! War or not, she’s determined to find a way to save her home and follow her dreams - whatever it takes.