When her family is struck by a terrible tragedy, Sarah Nightingale is forced to go into service at Grey Friars House as an under-nursery-maid. Although the situation wasn’t of her making Sarah thrives there, but life for her younger brother Alfie is not as easy. Desperate to avoid being sent to the workhouse, Alfie runs away to London to seek his fortune. But young and inexperienced with the dangerous streets of London, Alfie is tricked and sold to work as a slave on a coal barge. Sarah misses her brother terribly but knows she must find a way to survive if she ever wants them to be reunited.

When duty calls, East End girl Minnie Wolton isn’t afraid to do her bit. She’s excited to be joining the ATS girls and although she knows life at the barracks will be tough, she’s not one to quit! She just hopes the other girls are all good sorts. With her husband away fighting, Eileen Ruffel is glad of the break from her unhappy marriage. She just wants to make a success of the ATS girls and maybe find happiness too. Along with the other girls, Minnie and Eileen strike up a firm friendship. The training is hard, but they are determined to prove they are the best new recruits.

After Pearl's home is destroyed in the Exeter Blitz, so too are her dreams of performing onstage. Finding work as a bus conductress instead, a chance encounter revives her hopes once more, and soon she is singing for the troops alongside new friend Ivy. When agent Gordon Gold approaches them, Ivy jumps to sign with him and sets off for the bright lights of London. But Pearl is wary of the charming man and decides to stay, watching her friend go with a heavy heart. A year later, while Pearl is struck mute by an illness, Ivy returns - and is quick to seize the chance to fill Pearl's place.

Pearl has given up on her dreams of being a journalist after being denied a promotion because of her gender. Joining the WAAF as a radio telephone operator for Bomber Command, Pearl's new job at RAF Fenthorpe serves two purposes – providing an income for her grandmother, and keeping a close eye on her younger sister, Thea, an instrument repairer at another Lincolnshire bomber station.

Left homeless after the Exeter Blitz, May must find a job if she's to put a roof over her head and help support her two younger sisters. Taking a job as housekeeper to farmer George, May soon ends up getting stuck into more than just cleaning and cooking. The Ministry of Agriculture will close Fair Maids Farm if it doesn't meet their produce targets, but George refuses to heed their warnings. With only two reluctant Land Girls to help, May receives unexpected guidance from Dan, a neighbouring farmer, whose kindness gets tongues wagging in the village.