In 1944, as waves of German ninjas parachute into Kent, Britain's best hopes for victory lie with a Spitfire pilot codenamed Ack-Ack Macaque. The trouble is, Ack-Ack Macaque is a cynical, one-eyed, cigar-chomping monkey, and he's starting to doubt everything, including his own existence.
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In order to hide from his unwanted fame as the spitfire pilot-monkey who emerged from a computer game to defeat the nefarious corporation that engineered him, the charismatic and dangerous Ack-Ack Macaque is working as a pilot on a world-circling nuclear-powered zeppelin. But when the cabin of one of his passengers is invaded by the passenger's own dying doppelganger, our hirsute hero finds himself thrust into another race to save the world - this time from an aggressive hive mind, time-hopping saboteurs, and an army of homicidal Neanderthal assassins!
He's saved the world twice. Now the dangerous but charismatic Ack-Ack Macaque finds himself leading a dimension-hopping band of angry monkeys, facing an invading horde of implacable killer androids, and confronting the one challenge for which he was never prepared: impending fatherhood! Meanwhile, former journalist Victoria Valois finds herself facing old enemies as she fights to save the electronic ghost of her dead husband, and Merovech, king of the United Kingdoms of Great Britain and France, receives a troubling message from the dead sands of Mars....