Isla and her brother are two young foxes living just outside the lands of the furless - humans. The life of a fox is filled with dangers, but Isla has begun to learn mysterious skills meant to help her survive. Then the unthinkable happens. Returning to her den, Isla finds it set ablaze and surrounded by strange foxes, and her family is nowhere in sight. Forced to flee, she escapes into the cold, gray world of the furless. Now Isla must navigate this bewildering and deadly terrain, all while being hunted by a ruthless enemy.
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Isla's journey continues into the vast Wildlands as she seeks out her missing brother. As she travels, she finds her path is drawn inexorably toward the Elders, a group of wise and powerful foxes who have mastered the secrets of foxcraft.