When 10-year-old Megan helps her uncle invent the Thumbtop, the world's smallest computer, mice are overjoyed, and they want one for every mouse hole. The Big Cheese, leader of the Mouse Nation, has orders: Follow that girl - even if it means high-tailing it to Megan's new home on the other side of the country. While Megan struggles as the new girl, the mice wait for their chance.
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Life is good for the four humans in Cleveland who know that mice have evolved into possibly the smartest species on Earth. Eleven-year-old Megan and her cousin Joey have great summer jobs at the mouse-powered factory that makes Thumbtops - the tiny computers mice use to spread the word about climate change. But suddenly, the Big Cheese, leader of the Mouse Nation, needs their help. There's trouble at his Headquarters in California.
When a remote island's rain forest is threatened by the greedy timber company Loggocorp, the Humans Who Know (that mice have evolved) develop a secret plan to save the forest. And they need all the help they can get from the Big Cheese, Trey, and their other friends in the Mouse Nation. First the team must unravel a centuries-old mystery: Who actually owns the rain forest? The mission to find that person brings Megan and her family across the ocean to England, where everything starts to go just a little bit...weird.